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A dilute non-interacting suspension of micro-swimmers exhibits a finite velocity variance and short-ranged correlations that decay over a swimmer length. For a suspension of interacting straight swimmers, however, pair-interactions leads to…

Soft Condensed Matter · Physics 2020-12-02 Sankalp Nambiar , Piyush Garg , Ganesh Subramanian

Microswimmers typically operate in complex environments. In biological systems, often diverse species are simultaneously present and interact with each other. Here, we derive a (time-dependent) particle-scale statistical description, namely…

Soft Condensed Matter · Physics 2019-08-13 Christian Hoell , Hartmut Löwen , Andreas M. Menzel

In addition to enabling movement towards environments with favourable living conditions, swimming by microorganisms has also been linked to enhanced mixing and improved nutrient uptake by their populations. Experimental studies have shown…

Fluid Dynamics · Physics 2017-11-07 Blaise Delmotte , Eric E Keaveny , Eric Climent , Franck Plouraboué

In this paper, we explore the hydrodynamics of spheroidal active particles in viscosity gradients. This work provides a more accurate modeling approach, in comparison to spherical particles, for anisotropic organisms like Paramecium…

Fluid Dynamics · Physics 2025-07-09 Jiahao Gong , Vaseem A. Shaik , Gwynn J. Elfring

Particles transported in fluid flows, such as cells, polymers, or nanorods, are rarely spherical. In this study, we numerically and theoretically investigate the dispersion of an initially localized patch of passive elongated Brownian…

Fluid Dynamics · Physics 2021-09-15 Ajay Harishankar Kumar , Stuart J. Thomson , Thomas R. Powers , Daniel M. Harris

Previous particle-based computer simulations have revealed a significantly more pronounced tendency of spontaneous global polar ordering in puller (contractile) microswimmer suspensions than in pusher (extensile) suspensions. We here…

Soft Condensed Matter · Physics 2018-10-23 Christian Hoell , Hartmut Löwen , Andreas M. Menzel

In isotropic fluids like water, micrometer-scale swimmers have evolved swim strokes to translate despite their tiny size. As described by Purcell in his Scallop Theorem, reciprocal motions, like those performed by a scallop, cannot drive…

Mass dispersion in oscillatory flows is intimately linked to various environmental and biological processes, offering a distinct contrast to dispersion in steady flows due to the periodic expansion and contraction of particle patches. In…

Fluid Dynamics · Physics 2025-10-13 Bohan Wang , Weiquan Jiang , Li Zeng , Zi Wu , Ping Wang

The majority of studies on self-propelled particles and microswimmers concentrates on objects that do not feature a deterministic bending of their trajectory. However, perfect axial symmetry is hardly found in reality, and shape-asymmetric…

Statistical Mechanics · Physics 2018-07-18 Christian Hoell , Hartmut Löwen , Andreas M. Menzel

Microorganisms, such as E.Coli, are known to display upstream behavior and respond rheotactically to shear flows. In particular, E.Coli suspensions have been shown to display strong sensitivity to spatial constrictions, leading to an…

Soft Condensed Matter · Physics 2024-01-24 Armin Maleki , Malihe Ghodrat , Ignacio Pagonabarraga

We investigate how shear influences the dynamics of active particles in dilute to concentrated suspensions. Using apolar active suspensions of squirmers as model systems, we show how their long-time diffusive dynamics can surprisingly slow…

Soft Condensed Matter · Physics 2025-11-14 Zhouyang Ge , John F. Brady , Gwynn J. Elfring

Understanding the rheology of colloidal suspensions is crucial in the formulation of a wide selection of industry-relevant products. To characterise the viscoelastic behaviour of these soft materials, one can analyse the microscopic…

Soft Condensed Matter · Physics 2021-07-22 Fabián A. García Daza , Antonio M. Puertas , Alejandro Cuetos , Alessandro Patti

Marine organisms manipulate their surrounding flow through their swimming dynamics, which affects the transport of their own odor cues. We demonstrate by direct numerical simulations how a group of swimmers, moving at intermediate Reynolds…

Fluid Dynamics · Physics 2026-05-04 Martin James , Francesco Viola , Agnese Seminara

The rotational Brownian motion of colloidal spheres in dense suspensions reflects local hydrodynamics and friction, both key to non-linear rheological phenomena such as shear-thickening and jamming, and transport in crowded environments,…

Soft Condensed Matter · Physics 2021-06-23 Taiki Yanagishima , Yanyan Liu , Hajime Tanaka , Roel Dullens

This study uses a combination of stochastic optimization, statistical mechanical theory, and molecular simulation to test the extent to which the long-time dynamics of a single tracer particle can be enhanced by rationally modifying its…

Soft Condensed Matter · Physics 2011-11-24 James Carmer , Gaurav Goel , Mark J. Pond , Jeffrey R. Errington , Thomas M. Truskett

We investigate a model for the dynamics of ellipsoidal microswimmers in an externally imposed, laminar Kolmogorov flow. Through a phase-space analysis of the dynamics without noise, we find that swimmers favor either cross-stream or…

Soft Condensed Matter · Physics 2022-02-03 Simon A. Berman , Kyle S. Ferguson , Nathaniel Bizzak , Thomas H. Solomon , Kevin A. Mitchell

The dispersion of a passive colloid immersed in a bath of non-interacting and non-Brownian run-and-tumble microswimmers in two dimensions is analyzed using stochastic simulations and an asymptotic theory, both based on a minimal model of…

Soft Condensed Matter · Physics 2024-03-18 Tanumoy Dhar , David Saintillan

Various aspects of self-motility of chemically active colloids in Newtonian fluids can be captured by simple models for their chemical activity plus a phoretic slip hydrodynamic boundary condition on their surface. For particles of simple…

Soft Condensed Matter · Physics 2018-04-09 M. N. Popescu , W. E. Uspal , Z. Eskandari , M. Tasinkevych , S. Dietrich

Organisms often swim through density stratified fluids. In this Letter, we investigate the dynamics of small active particles swimming in density gradients and report theoretical evidence of taxis as a result of density stratification…

Soft Condensed Matter · Physics 2023-08-02 Vaseem A. Shaik , Gwynn J. Elfring

We experimentally and theoretically study the dynamics of a low-Reynolds number helical swimmer moving across viscosity gradients. Experimentally, a double-layer viscosity is generated by superposing two miscible fluids with similar…