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There is much current interest in modelling suspensions of algae and other micro-organisms for biotechnological exploitation, and many bioreactors are of tubular design. Using generalized Taylor dispersion theory, we develop a…

Biological Physics · Physics 2013-02-28 R. N. Bearon , M. A. Bees , O. A. Croze

Shear flow significantly affects the transport of swimming algae in suspension. For example, viscous and gravitational torques bias bottom-heavy cells to swim towards regions of downwelling fluid (gyrotaxis). It is necessary to understand…

Biological Physics · Physics 2015-03-20 O. A. Croze , G. Sardina , M. Ahmed , M. A. Bees , L. Brandt

The combined impact of axial stretching and cross-stream diffusion on the downstream transport of solute is termed Taylor dispersion. The dispersion of active suspensions is qualitatively distinct: viscous and external torques can establish…

Soft Condensed Matter · Physics 2025-05-27 Hakan Osman Caldag , Martin Alan Bees

Swimming microorganisms rely on effective mixing strategies to achieve efficient nutrient influx. Recent experiments, probing the mixing capability of unicellular biflagellates, revealed that passive tracer particles exhibit anomalous…

Soft Condensed Matter · Physics 2013-02-04 Irwin M. Zaid , Jörn Dunkel , Julia M. Yeomans

We study the dynamics of gyrotactic microswimmers suspended in homogeneous and isotropic turbulence by using direct numerical simulations (DNS). The swimmers are characterized by three non-dimensional parameters: their aspect ratio…

Soft Condensed Matter · Physics 2026-04-24 Suraj Kumar Nayak , Vishwanath Shukla , Akshay Bhatnagar

Classical Taylor-Aris dispersion theory is extended to describe the transport of suspensions of self-propelled dipolar cells in a tubular flow. General expressions for the mean drift and effective diffusivity are determined exactly in terms…

Soft Condensed Matter · Physics 2010-08-25 M. A. Bees , O. A. Croze

Ensembles of biological and artificial microswimmers produce long-range velocity fields with strong nonequilibrium fluctuations, which result in dramatic increase diffusivity of embedded particles (tracers). While such enhanced diffusivity…

Soft Condensed Matter · Physics 2019-02-20 Sergey Belan , Mehran Kardar

In the presence of a laminar shear flow, the diffusion of passive colloidal particles is enhanced in the direction parallel to the flow. This classical phenomenon is known as Taylor-Aris dispersion. Besides, microorganisms, such as active…

The experiments of Leptos et al. [Phys. Rev. Lett. 103, 198103 (2009)] show that the displacements of small particles affected by swimming microorganisms achieve a non-Gaussian distribution, which nevertheless scales diffusively -- the…

Soft Condensed Matter · Physics 2017-10-26 Jean-Luc Thiffeault

Active (i.e., self-propelled or swimming) particles moving through an isotropic fluid exhibit conventional diffusive behavior. We report anomalous diffusion of an active particle moving in an anisotropic, nematic background. Whilst the…

Soft Condensed Matter · Physics 2016-06-22 John Toner , Hartmut Löwen , Henricus H. Wensink

Mass dispersion in oscillatory flows is intimately linked to various environmental and biological processes, offering a distinct contrast to dispersion in steady flows due to the periodic expansion and contraction of particle patches. In…

Fluid Dynamics · Physics 2025-10-13 Bohan Wang , Weiquan Jiang , Li Zeng , Zi Wu , Ping Wang

Gyrotactic algae are bottom heavy, motile cells whose swimming direction is determined by a balance between a buoyancy torque directing them upwards and fluid velocity gradients. Gyrotaxis has, in recent years, become a paradigmatic model…

Chaotic Dynamics · Physics 2019-03-25 Massimo Cencini , Guido Boffetta , Matteo Borgnino , Filippo De Lillo

We study the dynamics and the statistics of dilute suspensions of gyrotactic swimmers, a model for many aquatic motile microorganisms. By means of extensive numerical simulations of the Navier-Stokes equations at different Reynolds numbers,…

Fluid Dynamics · Physics 2021-04-13 Matteo Borgnino , Guido Boffetta , Filippo De Lillo , Massimo Cencini

Swimming microorganisms can influence the diffusion of passive particles. The effect of this swimmer-particle interaction depends on different properties, such as the hydrodynamic field of the swimmer and the relative sizes of…

Soft Condensed Matter · Physics 2020-05-08 Florian von Rüling , Francine Kolley , Alexey Eremin

Transport phenomena of microswimmers in fluid flows play a crucial role in various biological processes, including bioconvection and cell sorting. In this paper, we investigate the dispersion behavior of chiral microswimmers in a simple…

Fluid Dynamics · Physics 2024-03-14 Keito Ogawa , Kenta Ishimoto

We study the small scale clustering of gyrotactic swimmers transported by a turbulent flow, when the intrinsic variability of the swimming parameters within the population is considered. By means of extensive numerical simulations, we find…

Fluid Dynamics · Physics 2017-02-22 Matteo Borgnino , Filippo De Lillo , Guido Boffetta

A suspension of gyrotactic microalgae Chlamydomonas augustae swimming in a cylindrical water vessel in solid-body rotation is studied. Our experiments show that swimming algae form an aggregate around the axis of rotation, whose intensity…

Fluid Dynamics · Physics 2015-11-05 M. Cencini , M. Franchino , F. Santamaria , G. Boffetta

The dispersion of a passive colloid immersed in a bath of non-interacting and non-Brownian run-and-tumble microswimmers in two dimensions is analyzed using stochastic simulations and an asymptotic theory, both based on a minimal model of…

Soft Condensed Matter · Physics 2024-03-18 Tanumoy Dhar , David Saintillan

Populations of swimming microorganisms produce fluid motions that lead to dramatically enhanced diffusion of tracer particles. Using simulations of suspensions of swimming particles in a periodic domain, we capture this effect and show that…

Soft Condensed Matter · Physics 2008-07-22 Patrick T. Underhill , Juan P. Hernández-Ortiz , Michael D. Graham

Living microorganisms are capable of a tactic response to external stimuli by swimming towards or away from the stimulus source; they do so by adapting their tactic signal transduction pathways to the environment. Their self-motility thus…

Soft Condensed Matter · Physics 2016-10-12 Alexander Geiseler , Peter Hänggi , Fabio Marchesoni , Colm Mulhern , Sergey Savel'ev
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