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The emergence of long-range spatiotemporal order from intrinsic chaos is a central challenge in far-from-equilibrium physics. In active fluids, such as cytoskeletal networks driving cellular motion, self-generated flows typically produce…

We use a continuum, two-fluid approach to study a mixture of two active nematic fluids. Even in the absence of thermodynamically-driven ordering, for mixtures of different activities we observe turbulent microphase separation, where domains…

Soft Condensed Matter · Physics 2025-02-03 Saraswat Bhattacharyya , Julia M. Yeomans

Active matter is a new class of material, intrinsically out-of equilibrium with intriguing properties. So far, the recent upsurge of studies has mostly focused on the spontaneous behavior of these systems --in the absence of external…

Soft Condensed Matter · Physics 2016-09-28 Nicolas Waisbord , Christopher Lefevre , Lyderic Bocquet , Christophe Ybert , Cecile Cottin-Bizonne

Bacterial cellulose is an important class of biomaterials which can be grown in well-controlled laboratory and industrial conditions. The cellulose structure is affected by several biological, chemical and environmental factors, including…

Biological Physics · Physics 2021-07-28 Sung-Ha Hong , Jia Yang , Mahdi Davoodianidalik , Horst Punzmann , Michael Shats , Hua Xia

Activity and renewability are distinctive features of living matter, and constitute a new class of materials that we term renewable active matter. A striking example is the cell cytoskeleton, where myosin filaments bind to the actin…

Soft Condensed Matter · Physics 2025-05-27 M. Abhishek , Ankit Dhanuka , Deb Sankar Banerjee , Madan Rao

Several types of active agents self-propel by spinning around their propulsion axis, thus behaving as active screws. Examples include cytoskeletal filaments in gliding assays, magnetically-driven colloidal helices, and microorganisms like…

Soft Condensed Matter · Physics 2026-02-03 Debarghya Banerjee , Lauritz Hahn , Ricard Alert

Microswimmers exhibit an intriguing, highly-dynamic collective motion with large-scale swirling and streaming patterns, denoted as active turbulence -- reminiscent of classical high-Reynolds-number hydrodynamic turbulence. Various…

Soft Condensed Matter · Physics 2024-06-03 Kai Qi , Elmar Westphal , Gerhard Gompper , Roland G. Winkler

The study of active matter system has critical importance in revealing the physical essence of biological collective behavior. Dense bacterial suspension - a typical biological active matter, exhibits a wide range of phenomenons, among…

Soft Condensed Matter · Physics 2025-11-04 Hongyi Bian , Chunhe Li , Zixiang Lin , Jin Zhu , Weijie Chen , Gaojin Li , Yongxiang Huang , Zijie Qu

We numerically study two-dimensional active nematics with periodic activity patterning. For stripes of activity, we observe a transition from two-dimensional to one-dimensional active turbulence as the maximum active force and distance…

Soft Condensed Matter · Physics 2025-03-11 Cody D. Schimming , C. J. O. Reichhardt , C. Reichhardt

Active gels are a class of biologically-relevant material containing embedded agents that spontaneously generate forces acting on a sparse filament network. In vitro experiments of protein filaments and molecular motors have revealed a…

Soft Condensed Matter · Physics 2024-06-03 David A. Head , Gerhard Gompper , W. J. Briels

The fate of every eukaryotic cell subtly relies on the exceptional mechanical properties of microtubules. Despite significant efforts, understanding their unusual mechanics remains elusive. One persistent, unresolved mystery is the…

Soft Condensed Matter · Physics 2015-06-19 Falko Ziebert , Hervé Mohrbach , Igor M. Kulić

Motor proteins drive persistent motion and self-organisation of cytoskeletal filaments. However, state-of-the-art microscopy techniques and continuum modelling approaches focus on large length and time scales. Here, we perform…

Soft Condensed Matter · Physics 2024-06-03 Gerrit Vliegenthart , Arvind Ravichandran , Marisol Ripoll , Thorsten Auth , Gerhard Gompper

We show that dense active fluids comprising interacting particles with persistent self-propulsion are driven to a non-equilibrium steady state consisting of co-moving particles with co-aligned active forces. This velocity and force sorting…

Soft Condensed Matter · Physics 2025-09-03 Suman Dutta , Pinaki Chaudhuri , Madan Rao , Chandan Dasgupta

Living systems are capable of locomotion, reconfiguration, and replication. To perform these tasks, cells spatiotemporally coordinate the interactions of force-generating, "active" molecules that create and manipulate non-equilibrium…

Soft Condensed Matter · Physics 2019-08-28 Tyler D. Ross , Heun Jin Lee , Zijie Qu , Rachel A. Banks , Rob Phillips , Matt Thomson

We review recent advances in the design, synthesis, and modeling of active fluids. Active fluids have been at the center of many technological innovations and theoretical advances over the past two decades. Research on this new class of…

Soft Condensed Matter · Physics 2023-02-02 Ilham Essafri , Bappa Ghosh , Caroline Desgranges , Jerome Delhommelle

Inspired by the Turing mechanism for pattern formation, we propose a simple self-propelled particle model with short-ranged alignment and anti-alignment at larger distances. It is able to produce orientationally ordered states, periodic…

Soft Condensed Matter · Physics 2016-05-02 Robert Grossmann , Pawel Romanczuk , Markus Bär , Lutz Schimansky-Geier

Using novel micro-printing techniques, we develop a versatile experimental setup that allows us to study how lateral confinement tames the active flows and defect properties of the microtubule/kinesin active nematic system. We demonstrate…

The study of liquid crystals at equilibrium has led to fundamental insights into the nature of ordered materials, as well as to practical applications such as display technologies. Active nematics are a fundamentally different class of…

Soft Condensed Matter · Physics 2015-08-19 Stephen J. DeCamp , Gabriel S. Redner , Aparna Baskaran , Michael F. Hagan , Zvonimir Dogic

The cytoskeleton relies on diverse populations of motors, filaments, and binding proteins acting in concert to enable non-equilibrium processes ranging from mitosis to chemotaxis. Its versatile reconfigurability, programmed by interactions…

We study spatiotemporal chaos in two-dimensional dense active suspensions using a generalized hydrodynamic model. Increasing activity induces a structural transition marked by the formation of intense vortices and giant number fluctuations…

Fluid Dynamics · Physics 2026-03-13 Kirti Kashyap , Kolluru Venkata Kiran , Anupam Gupta