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We discuss a polymer model for the 3D organization of human chromosomes. A chromosome is represented by a string of beads, with each bead being "colored" according to 1D bioinformatic data (e.g., chromatin state, histone modification, GC…

Biological Physics · Physics 2020-10-15 C. A. Brackley , D. Michieletto , F. Mouvet , J. Johnson , S. Kelly , P. R. Cook , D. Marenduzzo

Three-dimensional (3D) chromatin structure is closely related to genome function, in particular transcription. However, the folding path of the chromatin fiber in the interphase nucleus is unknown. Here, we systematically measured the 3D…

The 3D organisation of the genome in interphase cells is not a randomly folded polymer. Rather, experiments show that chromosomes arrange into a network of 3D compartments that correlate with biological processes, such as transcription,…

Biological Physics · Physics 2018-11-06 Kumar Rajendra , Ludvig Lizana , Per Stenberg

The three-dimensional organization of chromatin, on the length scale of a few genes, is crucial in determining the functional state - accessibility and the amount of gene expression - of the chromatin. Recent advances in chromosome…

Soft Condensed Matter · Physics 2021-07-16 K. Kumari , B. Duenweg , R. Padinhateeri , J. Ravi Prakash

The three-dimensional organisation of chromosomes can be probed using methods such as Capture-C. However it is unclear how such population level data relates to the organisation within a single cell, and the mechanisms leading to the…

Quantitative Methods · Quantitative Biology 2016-01-13 Chris A Brackley , Jill M Brown , Dominic Waithe , Christian Babbs , James Davies , Jim R Hughes , Veronica J Buckle , Davide Marenduzzo

Live-cell imaging experiments have shown that the distal dynamics between enhancers and promoters are unexpectedly rapid and incompatible with standard polymer models. The discordance between the compact static chromatin organization and…

Biological Physics · Physics 2025-06-24 Guang Shi , Sucheol Shin , D. Thirumalai

Understanding the three-dimensional (3D) structure of the genome is essential for elucidating vital biological processes and their links to human disease. To determine how the genome folds within the nucleus, chromosome conformation capture…

Three-Dimensional (3D) chromatin interactions, such as enhancer-promoter interactions (EPIs), loops, Topologically Associating Domains (TADs), and A/B compartments play critical roles in a wide range of cellular processes by regulating gene…

Genomics · Quantitative Biology 2024-03-07 Brydon P. G. Wall , My Nguyen , J. Chuck Harrell , Mikhail G. Dozmorov

Characterizing the link between small-scale chromatin structure and large-scale chromosome folding during interphase is a prerequisite for understanding transcription. Yet, this link remains poorly investigated. Here, we introduce a simple…

Soft Condensed Matter · Physics 2016-09-28 Ana Maria Florescu , Pierre Therizols , Angelo Rosa

Chromosome organisation is increasingly recognised as an essential component of genome regulation, cell fate and cell health. Within the realm of transposable elements (TEs) however, the spatial information of how genomes are folded is…

Biological Physics · Physics 2019-11-01 Alexandros Bousios , Hans-Wilhelm Nuetzmann , Dorothy Buck , Davide Michieletto

Genome organization in eukaryotes during interphase stems from the delicate balance between non-random correlations present in the DNA polynucleotide linear sequence and the physico/chemical reactions which shape continuously the form and…

Genomics · Quantitative Biology 2020-05-15 Alessandra Merlotti , Angelo Rosa , Daniel Remondini

How long threadlike eukaryotic chromosomes fit tidily in the small volume of the nucleus without significant entanglement is just beginning to be understood, thanks to major advances in experimental techniques. Several polymer models, which…

Soft Condensed Matter · Physics 2024-10-03 D. Thirumalai , Guang Shi , Sucheol Shin , Changbong Hyeon

Recent experiments have been able to visualise chromosome organization in fast-growing E.coli cells. However, the mechanism underlying the spatio-temporal organization remains poorly understood. We propose that the DNA adopts a specific…

Soft Condensed Matter · Physics 2026-04-06 Shreerang Pande , Debarshi Mitra , Apratim Chatterji

Increasing evidence suggests that chromosome folding and genetic expression are intimately connected. For example, the co-expression of a large number of genes can benefit from their spatial co-localization in the cellular space.…

Soft Condensed Matter · Physics 2018-03-13 Guillaume Le Treut

Current models for the folding of the human genome see a hierarchy stretching down from chromosome territories, through A/B compartments and TADs (topologically-associating domains), to contact domains stabilized by cohesin and CTCF.…

Biological Physics · Physics 2020-10-02 Peter R. Cook , Davide Marenduzzo

The three dimensional structure of DNA in the nucleus (chromatin) plays an important role in many cellular processes. Recent experimental advances have led to high-throughput methods of capturing information about chromatin conformation on…

Genomics · Quantitative Biology 2015-11-05 Kevin Emmett , Benjamin Schweinhart , Raul Rabadan

It is widely believed that the folding of the chromosome in the nucleus has a major effect on genetic expression. For example co-regulated genes in several species have been shown to colocalize in space despite being far away on the DNA…

Quantitative Methods · Quantitative Biology 2020-04-29 Guillaume Le Treut , François Képès , Henri Orland

We employ the diffusing transcription factors model for numerical simulation of chromatin topology conformations and transcriptional processes of human chromatin. Simulations of a short chromatin filament reveal different possible pathways…

Soft Condensed Matter · Physics 2023-07-10 Massimiliano Semeraro , Giuseppe Negro , Antonio Suma , Giuseppe Gonnella , Davide Marenduzzo

We propose a polymer model for the dynamics of chromatin replication in three dimensional space. Our simulations indicate that both immobile and tracking replisomes may self-assemble during the process, reconciling previous apparently…

Biological Physics · Physics 2023-12-19 G. Forte , S. Buonomo , P. R. Cook , N. Gilbert , D. Marenduzzo , E. Orlandini

Magnetic tweezers are used to study the mechanical response under torsion of single nucleosome arrays reconstituted on tandem repeats of 5S positioning sequences. Regular arrays are extremely resilient and can reversibly accommodate a large…

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