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Unlike macroscopic swimmers, microswimmers operate in a low-Reynolds-number regime dominated by viscous forces. This paper investigates the controllability of a magnetic microswimmer composed of a spherical magnetic head and an elastic,…

Optimization and Control · Mathematics 2025-11-05 Lucas Palazzolo , Mickaël Binois , Laëtitia Giraldi

Swimming micro-organisms such as flagellated bacteria and sperm cells have fascinating locomotion capabilities. Inspired by their natural motion, there is an ongoing effort to develop artificial robotic nano-swimmers for potential in-body…

Fluid Dynamics · Physics 2023-05-31 Jithu Paul , Yizhar Or , Oleg Gendelman

Microswimming cells and robots exhibit diverse behaviours due to both their swimming and their environment. One of the core environmental features impacting inertialess swimming is background flows. While the influence of select flows,…

Fluid Dynamics · Physics 2025-10-30 Eamonn A. Gaffney , Kenta Ishimoto , Benjamin J. Walker

An artificial microswimmer drifts in response to spatio-temporal modulations of an activating suspension medium. We consider two competing mechanisms capable of influencing its tactic response: angular fluctuations, which help it explore…

Soft Condensed Matter · Physics 2017-02-15 Alexander Geiseler , Peter Hänggi , Fabio Marchesoni

In a fluid environment, flagellated microswimmers propel themselves by rotating their flagella. The morphology of these flagella significantly influences forward speed, swimming efficiency, and directional stability, which are critical for…

Fluid Dynamics · Physics 2025-06-25 Baopi Liu , Lu Chen , Wenjun Xu

Systems of motile microorganisms exhibit a multitude of collective phenomena, including motility-induced phase separation and turbulence. Sensing of the environment and adaptation of movement plays an essential role in the emergent…

Soft Condensed Matter · Physics 2024-06-26 Segun Goh , Elmar Westphal , Roland G. Winkler , Gerhard Gompper

Guiding active microswimmers by external fields to requested target locations is a promising strategy to realize complex transport on the microscale. To this end, one possibility consists of attaching the microswimmers to orientable passive…

Fluid Dynamics · Physics 2020-02-19 Abdallah Daddi-Moussa-Ider , Maciej Lisicki , Hartmut Löwen , Andreas M. Menzel

Many biological microswimmers locomote by periodically beating the densely-packed cilia on their cell surface in a wave-like fashion. While the swimming mechanisms of ciliated microswimmers have been extensively studied both from the…

Soft Condensed Matter · Physics 2023-06-22 Hanliang Guo , Hai Zhu , Ruowen Liu , Marc Bonnet , Shravan Veerapaneni

In a low Reynolds number fluid environment that microswimmers encounter, back-and-forth motion cannot lead to net displacement. In mammalian sperm, the mechanical wave propagating along their single flagellum breaks the cancellation between…

Many biological microswimmers can modulate their swimming gait to achieve directional control of motility, especially when performing steering towards specific directional cues. This can be achieved without the need for obvious…

We study the three-dimensional dynamics of a spherical microswimmer in cylindrical Poiseuille flow which can be mapped onto a Hamiltonian system. Swinging and tumbling trajectories are identified. In 2D they are equivalent to oscillating…

Soft Condensed Matter · Physics 2012-06-18 Andreas Zöttl , Holger Stark

Synthetic microswimmers mimicking biological movements at the microscale have been developed in recent years. Actuating helical magnetic materials with a homogeneous rotating magnetic field is one of the most widespread techniques for…

Soft Condensed Matter · Physics 2019-03-27 Felix Bachmann Klaas Bente , Agnese Codutti , Damien Faivre

Microswimmers are active particles of microscopic size that self-propel by setting the surrounding fluid into motion. According to the kind of far-field fluid flow that they induce, they are classified into pushers and pullers. Many studies…

Soft Condensed Matter · Physics 2019-08-01 Giorgio Pessot , Hartmut Löwen , Andreas M. Menzel

For natural microswimmers, the interplay of swimming activity and external flow can promote robust motion, e.g. propulsion against ("upstream rheotaxis") or perpendicular to the direction of flow. These effects are generally attributed to…

Soft Condensed Matter · Physics 2018-01-30 Jaideep Katuri , William E. Uspal , Juliane Simmchen , Albert Miguel-López , Samuel Sánchez

When swimming at low Reynolds numbers, inertial effects are negligible and reciprocal movements cannot induce net motion. Instead, symmetry breaking is necessary to achieve net propulsion. Directed swimming can be supported by magnetic…

Soft Condensed Matter · Physics 2026-03-31 Theo Lequy , Andreas M. Menzel

Biological microswimmers are known to navigate upstream of an external flow (positive rheotaxis) in trajectories ranging from linear, spiral to oscillatory. Such rheotaxis stems from the interplay between the motion and complex shapes of…

Soft Condensed Matter · Physics 2022-05-27 Ranabir Dey , Carola M. Buness , Babak Vajdi Hokmabad , Chenyu Jin , Corinna C. Maass

Both natural and artificial small-scale swimmers may often self-propel in environments subject to complex geometrical constraints. While most past theoretical work on low-Reynolds number locomotion addressed idealised geometrical…

Fluid Dynamics · Physics 2017-11-16 Alexander Chamolly , Takuji Ishikawa , Eric Lauga

Self-propulsion of cellular microswimmers generates flow signatures, commonly classified as pusher- and puller-type, which characterize hydrodynamic interactions with other cells or boundaries. Using experimentally measured beat patterns,…

Cell Behavior · Quantitative Biology 2016-01-20 Gary S. Klindt , Benjamin M. Friedrich

We formulate and solve the equations governing the dynamics of a microscopic artificial swimmer composed of a head and of a tail made of a thin film of permanent magnetic material. This is a variant of the model swimmer proposed by Dreyfus…

Soft Condensed Matter · Physics 2017-02-15 Francois Alouges , Antonio DeSimone , Laetitia Giraldi , Marta Zoppello

The motion of microswimmers in complex flows is ruled by the interplay between swimmer propulsion and the dynamics induced by the fluid velocity field. Here we study the motion of a chiral microswimmer whose propulsion is provided by the…

Fluid Dynamics · Physics 2021-06-08 Ji Zhang , Mauro Chinappi , Luca Biferale
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