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Purcell's scallop theorem states that swimmers deforming their shapes in a time-reversible manner ("reciprocal" motion) cannot swim. Using numerical simulations and theoretical calculations we show here that in a fluctuating environment,…

Soft Condensed Matter · Physics 2011-08-30 Eric Lauga

The quest for the optimal navigation strategy in a complex environment is at the heart of microswimmer applications like cargo carriage or drug targeting to cancer cells. Here, we formulate a variational Fermat's principle for microswimmers…

Soft Condensed Matter · Physics 2019-01-25 Benno Liebchen , Hartmut Löwen

The possibility of microscopic swimming by extraction of energy from an external flow is discussed, focusing on the migration of a simple trimer across a linear shear flow. The geometric properties of swimming, together with the possible…

Soft Condensed Matter · Physics 2015-05-18 Piero Olla

The locomotion of microorganisms and spermatozoa in complex viscoelastic fluids is of critical importance in many biological processes such as fertilization, infection, and biofilm formation. Depending on their propulsion mechanisms,…

Soft Condensed Matter · Physics 2021-09-14 Gaojin Li , Eric Lauga , Arezoo M. Ardekani

We investigate the hydrodynamic interactions between microorganisms swimming at low Reynolds number. By considering simple model swimmers, and combining analytic and numerical approaches, we investigate the time-averaged flow field around a…

Soft Condensed Matter · Physics 2007-05-25 C. M. Pooley , G. P. Alexander , J. M. Yeomans

The swimming of an elliptical disk at small Reynolds number is studied on the basis of a perturbative solution of the Navier-Stokes equations for fluid flow near a deformable infinite sheet. A stroke involving an elliptically polarized…

Fluid Dynamics · Physics 2016-12-21 B. U. Felderhof

A stiff one-armed swimmer in glycerine goes nowhere, but if its arm is elastic, exerting a restorative torque proportional to local curvature, the swimmer can go on its way. Considering this happy consequence and the principles of…

Soft Condensed Matter · Physics 2009-10-30 Chris H. Wiggins , Raymond E. Goldstein

We present an automated procedure for the design of optimal actuation for flagellar magnetic microswimmers based on numerical optimization. Using this method, a new magnetic actuation method is provided which allows these devices to swim…

Soft Condensed Matter · Physics 2020-04-22 Yacine El Alaoui-Faris , Jean-Baptiste Pomet , Stéphane Régnier , Laetitia Giraldi

Efficient locomotion is important for the evolution of complex life, yet the physical principles selecting specific swimming strokes often remain entangled with biological constraints. In viscous fluids, the scallop theorem constrains the…

Biological Physics · Physics 2026-03-11 Takahiro Kanazawa , Kenta Ishimoto , Kyogo Kawaguchi

Here we introduce a two-dimensional (2D) low-Reynolds swimmer and discuss the motion of the swimmer both in noise-free and stochastic regimes. Three spheres, linked by extensible arms, in a plane form the triangle body of micro-swimmer.…

Soft Condensed Matter · Physics 2014-12-11 Mehran Ebrahimian , Mohammad Reza Ejtehadi

Synthetic microswimmers mimicking biological movements at the microscale have been developed in recent years. Actuating helical magnetic materials with a homogeneous rotating magnetic field is one of the most widespread techniques for…

Soft Condensed Matter · Physics 2019-03-27 Felix Bachmann Klaas Bente , Agnese Codutti , Damien Faivre

Large displacement optical flow is an integral part of many computer vision tasks. Variational optical flow techniques based on a coarse-to-fine scheme interpolate sparse matches and locally optimize an energy model conditioned on colour,…

Computer Vision and Pattern Recognition · Computer Science 2022-06-28 Qiao Chen , Charalambos Poullis

We study the distribution of swimming micro-organisms advected by a model turbulent flow and attracted towards a localised light source through phototaxis. It is shown that particles aggregate along a dynamical attractor with fractal…

Mathematical Physics · Physics 2009-11-13 Colin Torney , Zoltan Neufeld

We describe a method to track particles undergoing large displacements. Starting with a list of particle positions sampled at different time points, we assign particle identities by minimizing the sum across all particles of the trace of…

Soft Condensed Matter · Physics 2016-09-02 Rostislav Boltyanskiy , Jason W. Merrill , Eric R. Dufresne

Empirical observations show that turbulence exhibits a broad range of scaling exponents, characterizing how the velocity gradients diverge in the inviscid limit. These exponents are thought to be linked to singular solutions of the Euler…

Chaotic Dynamics · Physics 2025-11-11 Guillaume Costa , Amaury Barral , Adrien Lopez , Quentin Pikeroen , Bérengère Dubrulle

We study the dynamics of gyrotactic microswimmers suspended in homogeneous and isotropic turbulence by using direct numerical simulations (DNS). The swimmers are characterized by three non-dimensional parameters: their aspect ratio…

Soft Condensed Matter · Physics 2026-04-24 Suraj Kumar Nayak , Vishwanath Shukla , Akshay Bhatnagar

The swimming of a sphere immersed in a viscous incompressible fluid with inertia is studied for surface modulations of small amplitude on the basis of the Navier-Stokes equations. The mean swimming velocity and the mean rate of dissipation…

Fluid Dynamics · Physics 2016-12-20 B. U. Felderhof , R. B. Jones

The transport of bio-particles in viscous flows exhibits a rich variety of dynamical behaviour, such as morphological transitions, complex orientation dynamics or deformations. Characterising such complex behaviour under well controlled…

We study self propelled stokesian robots composed of assemblies of balls, in dimensions 2 and 3, and prove that they are able to control their position and orientation. This is a result of controllability, and its proof relies on applying…

Optimization and Control · Mathematics 2015-02-12 François Alouges , Antonio DeSimone , Luca Heltai , Aline Lefebvre , Benoît Merlet

Biological and artificial microswimmers often self-propel in external flows of vortical nature; relevant examples include algae in small-scale ocean eddies, spermatozoa in uterine peristaltic flows and bacteria in microfluidic devices. A…

Biological Physics · Physics 2022-11-14 Ivan Tanasijevic , Eric Lauga