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Flight is a complicated task at small scales in part due to the ubiquitous unsteady air which contains it. Flying organisms deal with these difficulties using active and passive control mechanisms to steer their body motion. Body attitudes…

Flapping insects are remarkably agile fliers, adapted to a highly turbulent environment. We present a series of high resolution numerical simulations of a bumblebee interacting with turbulent inflow. We consider both tethered and free…

Insects excel in trajectory and attitude handling during flight, yet the specific kinematic behaviours they use for maintaining stability in air disturbances are not fully understood. This study investigates the adaptive strategies of…

Quantitative Methods · Quantitative Biology 2024-09-04 Tim Jakobi , Simon Watkins , Alex Fisher , Sridhar Ravi

High-resolution numerical simulations of a tethered model bumblebee in forward flight are performed superimposing homogeneous isotropic turbulent fluctuations to the uniform inflow. Despite tremendous variation in turbulence intensity,…

Fluid Dynamics · Physics 2016-01-27 T. Engels , D. Kolomenskiy , K. Schneider , F. -O. Lehmann , J. Sesterhenn

In windy conditions, the air is turbulent. The strong and intermittent velocity variations of turbulence are invisible to flying animals. Nevertheless, flying animals, not much larger than the smallest scales of turbulence, manage to…

Fluid Dynamics · Physics 2023-04-25 Bardia Hejazi , Christian Küchler , Gholamhossein Bagheri , Eberhard Bodenschatz

The aerial environment in the operating domain of small-scale natural and artificial flapping wing fliers is highly complex, unsteady and generally turbulent. Considering flapping flight in an unsteady wind environment with a periodically…

Compared with fixed-wing flight, flapping flight can generate a higher lift and is also more maneuverable, largely resulting from the benefits of wing rotation. By analyzing the real wing kinematics of fruit flies, we found that the wing…

Fluid Dynamics · Physics 2019-06-19 Yang Xiang , Haotian Hang , Hong Liu

Birds rely on active high-acceleration morphing and flapping to navigate complex airflows, but they can also maintain stable fixed-wing postures under persistent atmospheric disturbances. Here, we show that avian wings exhibit aerodynamic…

Fluid Dynamics · Physics 2025-07-22 Lunbing Chen , Suyang Qin , Jinpeng Huang , Yufei Yin , Yang Xiang , Hong Liu

Hummingbirds and insects achieve outstanding flight performance by adapting their flapping motion to the flight requirements. Their wing kinematics can change from smooth flapping to highly dynamic waveforms, generating unsteady aerodynamic…

Fluid Dynamics · Physics 2024-08-07 Romain Poletti , Andre Calado , Lilla K. Koloszar , Joris Degroote , Miguel A. Mendez

High resolution direct numerical simulations of rotating and flapping bumblebee wings are presented and their aerodynamics is studied focusing on the role of leading edge vortices and the associated helicity production. We first study the…

Fluid Dynamics · Physics 2018-03-21 T. Engels , D. Kolomenskiy , K. Schneider , M. Farge , F. -O. Lehmann , J. Sesterhenn

Turbulence is omnipresent in the atmosphere and a long-standing scientific conundrum that makes flight complex. This complexity is little understood; surprisingly, when turbulence arises, air vehicles struggle while birds seem to thrive.…

Fluid Dynamics · Physics 2026-01-01 Ariane Gayout , David Lentink

Turbulent winds and gusts fluctuate on a wide range of timescales from milliseconds to minutes and longer, a range that overlaps the timescales of avian flight behavior, yet the importance of turbulence to avian behavior is unclear. By…

Quantitative Methods · Quantitative Biology 2021-07-07 Kasey M. Laurent , Bob Fogg , Tobias Ginsburg , Casey Halverson , Michael Lanzone , Tricia A. Miller , David W. Winkler , Gregory P. Bewley

Bio-inspired methods can provide efficient solutions to perform autonomous landing for Micro Air Vehicles (MAVs). Flying insects such as honeybees perform vertical landings by keeping flow divergence constant. This leads to an exponential…

Robotics · Computer Science 2016-09-23 H. W. Ho , G. C. H. E. de Croon , E. van Kampen , Q. P. Chu , M. Mulder

Wing flexibility plays an essential role in the aerodynamic performance of insects due to the considerable deformation of their wings during flight under the impact of inertial and aerodynamic forces. These forces come from the complex wing…

Fluid Dynamics · Physics 2020-07-20 Hung Truong , Thomas Engels , Dmitry Kolomenskiy , Kai Schneider

Insects and birds are often faced by opposing requirements for agile and stable flight. Here, we explore the interplay between aerodynamic effort, maneuverability, and stability in a model system that consists of a $\Lambda$-shaped flyer…

Fluid Dynamics · Physics 2014-11-26 Yangyang Huang , Monika Nitsche , Eva Kanso

Saving energy and enhancing performance are secular preoccupations shared by both nature and human beings. In animal locomotion, flapping flyers or swimmers rely on the flexibility of their wings or body to passively increase their…

Biological Physics · Physics 2011-08-30 Sophie Ramananarivo , Ramiro Godoy-Diana , Benjamin Thiria

Insects use flight muscles attached at the base of the wings to produce impressive wing flapping frequencies. The maximum power output of these flight muscles is insufficient to maintain such wing oscillations unless there is good elastic…

Fluid Dynamics · Physics 2016-07-20 Yangyang Huang , Eva Kanso

Aeroelastic structures, from insect wings to wind turbine blades, experience transient unsteady aerodynamic loads that are coupled to their motion. Effective real-time control of flexible structures relies on accurate and efficient…

Fluid Dynamics · Physics 2022-07-14 Michelle Hickner , Urban Fasel , Aditya G. Nair , Bingni W. Brunton , Steven L. Brunton

The movement of organisms is subject to a multitude of influences of widely varying character: from the bio-mechanics of the individual, over the interaction with the complex environment many animals live in, to evolutionary pressure and…

Biological Physics · Physics 2013-07-18 Friedrich Lenz , Aleksei V. Chechkin , Rainer Klages

Wing flapping is one of the most widespread propulsion methods found in nature; however, the current understanding of the aerodynamics in bird wakes is incomplete. The role of the unsteady motion in the flow and its contribution to the…

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