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Tanglegrams are a special class of graphs appearing in applications concerning cospeciation and coevolution in biology and computer science. They are formed by identifying the leaves of two rooted binary trees. We give an explicit formula…

Combinatorics · Mathematics 2015-07-20 Sara Billey , Matjaž Konvalinka , Frederick A Matsen

Rooted binary perfect phylogenies provide a generalization of rooted binary unlabeled trees in which each leaf is assigned a positive integer value that corresponds in a biological setting to the count of the number of indistinguishable…

Populations and Evolution · Quantitative Biology 2024-10-22 Chloe E. Shiff , Noah A. Rosenberg

Phylogenetic (i.e. leaf-labeled) trees play a fundamental role in evolutionary research. A typical problem is to reconstruct such trees from data like DNA alignments (whose columns are often referred to as characters), and a simple…

Populations and Evolution · Quantitative Biology 2022-09-08 Mareike Fischer

We investigate the number of permutations that occur in random labellings of trees. This is a generalisation of the number of subpermutations occurring in a random permutation. It also generalises some recent results on the number of…

Probability · Mathematics 2022-12-22 Michael Albert , Cecilia Holmgren , Tony Johansson , Fiona Skerman

Phylogenetic trees play a key role in the reconstruction of evolutionary relationships. Typically, they are derived from aligned sequence data (like DNA, RNA, or proteins) by using optimization criteria like, e.g., maximum parsimony (MP).…

Populations and Evolution · Quantitative Biology 2025-06-11 Mirko Wilde , Mareike Fischer

Null models of binary phylogenetic trees are useful for testing hypotheses on real world phylogenies. In this paper we consider phylogenies as binary trees without edge lengths together with a sampling measure and encode them as algebraic…

Probability · Mathematics 2020-06-17 Josué Nussbaumer , Anita Winter

A phylogenetic tree is an acyclic graph with distinctly labeled leaves, whose internal edges have a positive weight. Given a set of n leaves, the collection of all phylogenetic trees with this leaf set can be assembled into a metric cube…

Combinatorics · Mathematics 2019-04-02 Gillian Grindstaff

In 1989 Erd\H{o}s and Sz\'ekely showed that there is a bijection between (i) the set of rooted trees with $n+1$ vertices whose leaves are bijectively labeled with the elements of $[\ell]=\{1,2,\dots,\ell\}$ for some $\ell \leq n$, and (ii)…

Discrete Mathematics · Computer Science 2025-10-29 Vincent Moulton , Andreas Spillner

Phylogenetic trees are frequently used to model evolution. Such trees are typically reconstructed from data like DNA, RNA, or protein alignments using methods based on criteria like maximum parsimony (amongst others). Maximum parsimony has…

Populations and Evolution · Quantitative Biology 2023-07-31 Mirko Wilde , Mareike Fischer

A fringe subtree of a rooted tree is a subtree consisting of one of the nodes and all its descendants. In this paper, we are specifically interested in the number of non-isomorphic trees that appear in the collection of all fringe subtrees…

Combinatorics · Mathematics 2020-03-09 Louisa Seelbach Benkner , Stephan Wagner

Cayley's formula states that the number of labelled trees on $n$ vertices is $n^{n-2}$, and many of the current proofs involve complex structures or rigorous computation. We present a bijective proof of the formula by providing an…

Combinatorics · Mathematics 2014-09-08 Steven Hao , Andrew He , Ray Li , Scott Wu

Tanglegrams are drawings of two rooted binary phylogenetic trees and a matching between their leaf sets. The trees are drawn crossing-free on opposite sides with their leaf sets facing each other on two vertical lines. Instead of minimizing…

Data Structures and Algorithms · Computer Science 2023-05-09 Alexander Dobler , Martin Nöllenburg

A tanglegram is a pair of binary trees with the same set of leaves. Unlabeled tanglegrams were counted recently by Billey, Konvalinka, and Matsen, who also proposed the problem of counting several variations of unlabeled tanglegrams…

Combinatorics · Mathematics 2021-07-06 Ira M. Gessel

There are several common ways to encode a tree as a matrix, such as the adjacency matrix, the Laplacian matrix (that is, the infinitesimal generator of the natural random walk), and the matrix of pairwise distances between leaves. Such…

Populations and Evolution · Quantitative Biology 2007-05-23 Frederick A. Matsen , Steven N. Evans

We consider the rooted trees which not have isomorphic representation and introduce a conception of complexity a natural number also. The connection between quantity such trees with $n$ edges and a complexity of natural number $n$ is…

Combinatorics · Mathematics 2012-05-03 B. S. Kochkarev

In this article, we construct a generalization of the Blum-Fran\c{c}ois Beta-splitting model for evolutionary trees, which was itself inspired by Aldous' Beta-splitting model on cladograms. The novelty of our approach allows for asymmetric…

Probability · Mathematics 2016-07-04 Raazesh Sainudiin , Amandine Veber

Nested (or reconciled) phylogenetic trees model co-evolutionary systems in which one evolutionary history is embedded within another. We introduce a geometric framework for such systems by defining $\sigma$-space, a moduli space of fully…

Combinatorics · Mathematics 2026-04-16 G. Grindstaff , R. S. Hoekzema

We study the average number of distinct fringe subtrees in random trees generated by leaf-centric binary tree sources as introduced by Zhang, Yang and Kieffer. A leaf-centric binary tree source induces for every $n \geq 2$ a probability…

Discrete Mathematics · Computer Science 2025-04-30 Louisa Seelbach Benkner , Markus Lohrey , Stephan Wagner

As part of work to connect phylogenetics with machine learning, there has been considerable recent interest in vector encodings of phylogenetic trees. We present a simple new "ordered leaf attachment" (OLA) method for uniquely encoding a…

Populations and Evolution · Quantitative Biology 2025-03-26 Harry Richman , Cheng Zhang , Frederick A. Matsen

For a labelled tree on the vertex set $[n]:=\{1,2,..., n\}$, define the direction of each edge $ij$ to be $i\to j$ if $i<j$. The indegree sequence of $T$ can be considered as a partition $\lambda \vdash n-1$. The enumeration of trees with a…

Combinatorics · Mathematics 2009-04-02 Rosena R. X. Du , Jingbin Yin
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