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Fluid mechanics plays a vital role in early vertebrate embryo development, an example being the establishment of left-right asymmetry. Following the dorsal-ventral and anterior-posterior axes, the left-right axis is the last to be…

Fluid Dynamics · Physics 2013-09-06 Andrew A. Smith , Thomas D. Johnson , David J. Smith , John R. Blake

In embryonic development, programmed cell shape changes are essential for building functional organs, but in many cases the mechanisms that precisely regulate these changes remain unknown. We propose that fluid-like drag forces generated by…

Biological Physics · Physics 2019-01-03 Gonca Erdemci-Tandogan , Madeline J. Clark , Jeffrey D. Amack , M. Lisa Manning

Left-right symmetry breaking is critical to vertebrate embryonic development; in many species this process begins with cilia-driven flow in a structure termed the `node'. Primary `whirling' cilia, tilted towards the posterior, transport…

Fluid Dynamics · Physics 2010-07-13 David J. Smith , Andrew A. Smith , John R. Blake

Cells or bacteria carrying cilia on their surface show many striking features : alignment of cilia in an array, two-phase asymmetric beating for each cilium, coordination between cilia and existence of metachronal waves with a constant…

Biological Physics · Physics 2009-11-11 Boris Guirao , Jean-François Joanny

The concept of internal anatomical asymmetry is familiar; usually in humans the heart is on the left and the liver is on the right, however how does the developing embryo know to produce this consistent laterality? Symmetry breaking…

Tissues and Organs · Quantitative Biology 2019-10-10 Meurig T. Gallagher , Thomas D. Montenegro-Johnson , David J. Smith

Cilia and flagella are actively bending slender organelles, performing functions such as motility, feeding and embryonic symmetry breaking. We review the mechanics of viscous-dominated microscale flow, including time-reversal symmetry, drag…

Quantitative Methods · Quantitative Biology 2013-09-06 Thomas D. Montenegro-Johnson , Andrew A. Smith , David J. Smith , Daniel Loghin , John R. Blake

The dynamic flow of cerebrospinal fluid (CSF) in brain ventricles exhibits flow features on several scales, both spatially and temporally. Most analysis of this complex flow and the accompanying transport has used instantaneous (Eulerian)…

Fluid Dynamics · Physics 2026-03-20 Halvor Herlyng , Shawn C. Shadden

The proposed flow in a 3-D cubic cavity is driven by its parallel walls moving in perpendicular directions to create a genuinely three-dimensional highly separated vortical flow yet having simple single-block cubical geometry of…

Fluid Dynamics · Physics 2023-01-23 Alex Povitsky

In this paper, we showcase how flow obstruction by a deformable object can lead to symmetry breaking in curved domains subject to angular acceleration. Our analysis is motivated by the deflection of the cupula, a soft tissue located in the…

Fluid Dynamics · Physics 2025-11-06 Javier Chico-Vázquez , Derek E. Moulton , Dominic Vella

The laboratory models of the human heart left ventricle developed in the last decades gave a valuable contribution to the comprehension of the role of the fluid dynamics in the cardiac function and to support the interpretation of the data…

Fluid Dynamics · Physics 2014-07-29 S. Fortini , G. Querzoli , S. Espa , A. Cenedese

In the present paper, we examine the viscous flow evolution in a square cavity. Coupled with the stream function, the initial-boundary value problem of the vorticity is numerically solved by a method of iteration. The only boundary…

Fluid Dynamics · Physics 2018-04-12 F. Lam

Motile cilia beat in an asymmetric fashion in order to propel the surrounding fluid. When many cilia are located on a surface, their beating can synchronise such that their phases form metachronal waves. Here, we computationally study a…

Soft Condensed Matter · Physics 2023-09-29 David J. Hickey , Ramin Golestanian , Andrej Vilfan

To gain insight into the nature of biological synchronization at the microscopic scale, we here investigate the hydrodynamic synchronization between conically rotating objects termed nodal cilia. A mechanical model of three rotating cilia…

Soft Condensed Matter · Physics 2018-03-28 Keiji Okumura , Seiya Nishikawa , Toshihiro Omori , Takuji Ishikawa , Atsuko Takamatsu

The study numerically investigates the three-dimensionality in the flow and two-degree-of-freedom (2 DOF) vortex-induced-vibrations (VIV) characteristics of an elastically mounted circular cylinder. The cylinder is allowed to vibrate in…

Fluid Dynamics · Physics 2022-11-08 Mayank Verma , Ashoke De

We calculate the hydrodynamic flow field generated far from a cilium which is attached to a surface and beats periodically. In the case of two beating cilia, hydrodynamic interactions can lead to synchronization of the cilia, which are…

Biological Physics · Physics 2007-05-23 Andrej Vilfan , Frank Julicher

We use numerical simulations to systematically investigate the vesicle dynamics in two-dimensional (2D) Taylor-Green vortex flow in the absence of inertial forces. Vesicles are highly deformable membranes encapsulating an incompressible…

Biological Physics · Physics 2023-05-10 Gökberk Kabacaoğlu , Enkeleida Lushi

During embryonic development in vertebrates, left-right (L/R) asymmetry is reliably generated by a conserved mechanism: a L/R asymmetric signal is transmitted from the embryonic node to other parts of the embryo by the L/R asymmetric…

Tissues and Organs · Quantitative Biology 2011-12-16 Hanrong Chen , C. L. Henley , B. Xu

We introduce a model system of stochastic entities, called 'rowers' which include some essentialities of the behavior of real cilia. We introduce and discuss the problem of symmetry breaking for these objects and its connection with the…

Statistical Mechanics · Physics 2009-11-07 M. Cosentino Lagomarsino , B. Bassetti , P. Jona

In this work, the coupled dynamics of the gap flow and the vortex-induced vibration (VIV) on a side-by-side (SBS) arrangement of two circular cylinders is numerically investigated at moderate Reynolds number 100 < Re < 800. Of particular…

Fluid Dynamics · Physics 2023-07-19 B. Liu , R. K. Jaiman

Clockwise rotational movement of isolated single cilia in mice embryo was investigated in vivo. The movement generates leftward fluid flow in the node cavity and plays an important role in left-right determination. The leftward…

Tissues and Organs · Quantitative Biology 2013-05-22 Atsuko Takamatsu , Takuji Ishikawa , Kyosuke Shinohara , Hiroshi Hamada
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