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Related papers: Characteristics of undulatory locomotion in granul…

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Undulatory swimming is a widespread propulsion strategy adopted by many small-scale organisms including various single-cell eukaryotes and nematodes. In this work, we report a comprehensive study of undulatory locomotion of a finite…

Fluid Dynamics · Physics 2015-06-16 R. Berman , O. Kenneth , J. Sznitman , A. Leshansky

We use high-speed x-ray imaging to reveal how a small (~10cm) desert dwelling lizard, the sandfish (Scincus scincus), swims within a granular medium [1]. On the surface, the lizard uses a standard diagonal gait, but once below the surface,…

Fluid Dynamics · Physics 2009-10-20 Ryan D. Maladen , Yang Ding , Adam Kamor , Daniel I. Goldman

Sand is a highly dissipative system, where the local spatial arrangements and densities depend strongly on the applied forces, resulting in fluid-like or solid-like behaviour. This makes sand swimming challenging and intriguing, raising…

Soft Condensed Matter · Physics 2024-04-23 Inaki Echeverría-Huarte , Margarida M. Telo da Gama , Nuno A. M. Araujo

Undulatory locomotion, a gait in which thrust is produced in the opposite direction of a traveling wave of body bending, is a common mode of propulsion used by animals in fluids, on land, and even within sand. As such it has been an…

Biological Physics · Physics 2013-06-11 Yang Ding , Sarah S. Sharpe , Kurt Wiesenfeld , Daniel I. Goldman

Undulatory locomotion is a means of self-propulsion that relies on the generation and propagation of waves along a body. As a mode of locomotion it is primitive and relatively simple, yet can be remarkably robust. No wonder then, that it is…

Biological Physics · Physics 2009-08-20 Netta Cohen , Jordan H. Boyle

Many biological fluids are composed of suspended polymers immersed in a viscous fluid. A prime example is mucus, where the polymers are also known to form a network. While the presence of this microstructure is linked with an overall…

Fluid Dynamics · Physics 2024-10-10 Adam K. Townsend , Eric E. Keaveny

Although typically possessing four limbs and short bodies, lizards have evolved a diversity of body plans, from short-bodied and fully-limbed to elongate and nearly limbless. Such diversity in body morphology is hypothesized as adaptations…

Biological Physics · Physics 2022-08-10 Baxi Chong , Tianyu Wang , Eva Erickson , Philip J Bergmann , Daniel I. Goldman

Understanding swimming in soft yielding media is challenging due to their complex deformation response to the swimmer's motion. We experimentally show that a scallop-inspired swimmer with reciprocally flapping wings generates locomotion in…

Soft Condensed Matter · Physics 2024-12-09 Hongyi Xiao , Harol Torres , Achim Sack , Thorsten Pöschel

Swimming cells and microorganisms must often move though complex fluids that contain an immersed microstructure such as polymer molecules, or filaments. In many important biological processes, such as mammalian reproduction and bacterial…

Fluid Dynamics · Physics 2018-08-06 Arshad Kamal , Eric E Keaveny

In this theoretical study, we present an analytical framework to investigate the slithering motion of snakes on flat surfaces. While previous studies have predominantly relied on numerical methods to identify optimal locomotion kinematics,…

Biological Physics · Physics 2024-12-06 Yogesh Phalak , Hodjat Pendar

Many cells exploit the bending or rotation of flagellar filaments in order to self-propel in viscous fluids. While appropriate theoretical modelling is available to capture flagella locomotion in simple, Newtonian fluids, formidable…

Biological Physics · Physics 2017-08-02 Emily E. Riley , Eric Lauga

Micro-organisms expend energy moving through complex media. While propulsion speed is an important property of locomotion, efficiency is another factor that may determine the swimming gait adopted by a micro-organism in order to locomote in…

Fluid Dynamics · Physics 2017-12-14 Herve Nganguia , Kyle Pietrzyk , On Shun Pak

Motivated by the swimming of sperm in the non-Newtonian fluids of the female mammalian reproductive tract, we examine the swimming of filaments in the nonlinear viscoelastic Upper Convected Maxwell model. We obtain the swimming velocity and…

Soft Condensed Matter · Physics 2009-11-13 Henry C. Fu , Thomas R. Powers , Charles W. Wolgemuth

We study a simple model of periodic contraction and extension of large intruders in a granular bed to understand the mechanism for swimming in an otherwise solid media. Using an event-driven simulation, we find optimal conditions that…

Statistical Mechanics · Physics 2015-05-14 Takashi Shimada , Dirk Kadau , Troy Shinbrot , Hans J. Herrmann

The physical and bio-mechanical principles that govern undulatory movement on wet surfaces have important applications in physiology, physics, and engineering. The nematode {\it C. elegans}, with its highly stereotypical and functionally…

Fluid Dynamics · Physics 2015-06-03 Xiao N. Shen , J. Sznitman , P. Krajacic , T. Lamitina , P. E. Arratia

Biological locomotion in nature is often achieved by the interaction between a flexible body and its surrounding medium. The interaction of a flexible body with granular media is less understood compared with viscous fluids partially due to…

Fluid Dynamics · Physics 2017-08-03 Zhiwei Peng , Yang Ding , Kyle Pietrzyk , Gwynn J. Elfring , On Shun Pak

We use particle simulations to reveal two distinct propulsion mechanisms for a scallop-like swimmer to locomote itself in granular media by reciprocally flapping its wings. Based on the discrete element method, we examine the kinematics and…

Soft Condensed Matter · Physics 2025-10-28 Amir Nazemi , Hongyi Xiao

Motile eukaryotic cells propel themselves in viscous fluids by passing waves of bending deformation down their flagella. An infinitely long flagellum achieves a hydrodynamically optimal low-Reynolds number locomotion when the angle between…

Biological Physics · Physics 2010-03-03 Saverio E. Spagnolie , Eric Lauga

We investigate the dynamics of \textbf{\textit{Lumbriculus variegatus}} in water-saturated sediment beds to understand limbless locomotion in the benthic zone found at the bottom of lakes and oceans. These slender aquatic worms are observed…

Soft Condensed Matter · Physics 2020-11-30 Arshad Kudrolli , Bernny Ramirez

Self-propelling organisms locomote via generation of patterns of self-deformation. Despite the diversity of body plans, internal actuation schemes and environments in limbless vertebrates and invertebrates, such organisms often use similar…

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