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Many microorganisms swim through gels and non-Newtonian fluids in their natural environments. In this paper, we focus on microorganisms which use flagella for propulsion. We address how swimming velocities are affected in nonlinearly…

Biological Physics · Physics 2010-04-07 Henry C. Fu , Charles W. Wolgemuth , Thomas R. Powers

A number of swimming microorganisms such as ciliates ($\textit{Opalina}$) and multicellular colonies of flagellates ($\textit{Volvox}$) are approximately spherical in shape and swim using beating arrays of cilia or short flagella covering…

Fluid Dynamics · Physics 2014-02-06 On Shun Pak , Eric Lauga

When swimming at low Reynolds numbers, inertial effects are negligible and reciprocal movements cannot induce net motion. Instead, symmetry breaking is necessary to achieve net propulsion. Directed swimming can be supported by magnetic…

Soft Condensed Matter · Physics 2026-03-31 Theo Lequy , Andreas M. Menzel

Both natural and artificial small-scale swimmers may often self-propel in environments subject to complex geometrical constraints. While most past theoretical work on low-Reynolds number locomotion addressed idealised geometrical…

Fluid Dynamics · Physics 2017-11-16 Alexander Chamolly , Takuji Ishikawa , Eric Lauga

This paper studies the influence of orienting external fields on pattern formation, particularly mesoscale turbulence, in microswimmer suspensions. To this end, we apply a hydrodynamic theory that can be derived from a microscopic…

Soft Condensed Matter · Physics 2019-02-27 Henning Reinken , Sebastian Heidenreich , Markus Bär , Sabine H. L. Klapp

The swimming of cells, far from any boundary, can arise in the absence of long-range liquid-crystalline order within the cytoplasm, but simple models of this effect are lacking. Here we present a two-dimensional model of droplet…

Soft Condensed Matter · Physics 2020-07-29 Rajesh Singh , Elsen Tjhung , Michael E. Cates

A conformation space kinetic model is constructed to drive the deformation cycle of a three-sphere swimmer to achieve propulsion at low Reynolds number. We analyze the effect of an external load on the performance of this kinetic swimmer,…

Soft Condensed Matter · Physics 2009-11-13 Ramin Golestanian , Armand Ajdari

Hypothesis: Immiscible liquids are commonly used to achieve unique functions in many applications, where the breakup of compound droplets in airflow is an important process. Due to the existence of the liquid-liquid interface, compound…

Fluid Dynamics · Physics 2023-12-27 Zhikun Xu , Yue Zhang , Tianyou Wang , Zhizhao Che

We study the steady-state behavior of active, dipolar, Brownian spheroids in a planar channel subjected to an imposed Couette flow and an external transverse field, applied in the 'downward' normal-to-flow direction. The field-induced…

Soft Condensed Matter · Physics 2021-08-04 Mohammad Reza Shabanniya , Ali Naji

A fluid droplet in general deforms, if subject to active driving, such as a finite slip velocity or active tractions on its interface. We show that these deformations and their dynamics can be computed analytically in a perturbation theory…

Soft Condensed Matter · Physics 2023-04-19 Reiner Kree , Annette Zippelius

In isotropic fluids like water, micrometer-scale swimmers have evolved swim strokes to translate despite their tiny size. As described by Purcell in his Scallop Theorem, reciprocal motions, like those performed by a scallop, cannot drive…

Biological microswimmers alter their motility in complex corner geometries, facilitating their survival. However, the dynamical features of low-Reynolds-number swimming at corners remain undefined. Here, we use active droplet microswimmers…

Soft Condensed Matter · Physics 2026-04-22 Subhasish Guchhait , Harshita Tiwari , Sumesh P. Thampi , Ranabir Dey

We investigate the modes of deformation of an initially spherical bubble immersed in a homogeneous and isotropic turbulent background flow. We perform direct numerical simulations of the two-phase incompressible Navier-Stokes equations,…

Fluid Dynamics · Physics 2021-07-07 Stéphane Perrard , Aliénor Rivière , Wouter Mostert , Luc Deike

Micro-swimmer locomotion in heterogeneous media is increasingly relevant in biological physics due to the prevalence of microorganisms in complex environments. A model for such porous media is the Brinkman fluid which accounts for a sparse…

Fluid Dynamics · Physics 2026-02-05 Francisca Guzman-Lastra , Enkeleida Lushi

The dynamics of periodic swimming is studied for two models of a deformable sphere, the dipole-quadrupole model and the quadrupole-octupole model. For the two models the solution of the Navier-Stokes equations can be found exactly to second…

Fluid Dynamics · Physics 2019-12-18 B. U. Felderhof , R. B. Jones

The encapsulation of active particles, such as bacteria or active colloids, inside a droplet gives rise to nontrivial shape dynamics and droplet motility. To understand this behavior, we derive an asymptotic solution for the fluid flow…

Soft Condensed Matter · Physics 2025-03-19 Sho Kawakami , Petia M. Vlahovska

Undulatory slender objects have been a central theme in the hydrodynamics of swimming at low Reynolds number, where the slender body is usually assumed to be inextensible, although some microorganisms and artificial microrobots largely…

Fluid Dynamics · Physics 2025-11-10 Kenta Ishimoto , Johann Herault , Clément Moreau

The interaction of a droplet with a swirling airstream is investigated experimentally by shadowgraphy and particle image velocimetry techniques. In swirl flow, the droplet experiences oppose-flow, cross-flow, and co-flow conditions…

Fluid Dynamics · Physics 2023-06-05 Pavan Kumar Kirar , Surendra Kumar Soni , Pankaj S. Kolhe , Kirti Chandra Sahu

We report on measurements of the angular dynamics of micron-sized equilaterally triangular platelets suspended in a micro-channel shear flow. Our measurements confirm that such particles spin and tumble like a spheroid in a simple shear.…

Fluid Dynamics · Physics 2018-03-26 J. Fries , M. Vijay Kumar , B. Mekonnen Mihiretie , D. Hanstorp , B. Mehlig

Active systems comprised of self-propelled units show fascinating transitions from Brownian-like dynamics to collective coherent motion. Swirling of swimming bacteria is a spectacular example. This study demonstrates that a nematic liquid…

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