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From the formation of animal flocks to the emergence of coordinate motion in bacterial swarms, at all scales populations of motile organisms display coherent collective motion. This consistent behavior strongly contrasts with the difference…

Soft Condensed Matter · Physics 2013-11-11 Antoine Bricard , Jean-Baptiste Caussin , Nicolas Desreumaux , Olivier Dauchot , Denis Bartolo

The motion of active colloids is generally achieved through their anisotropy, as exemplified by Janus colloids. Recently, there was a growing interest in the propulsion of isotropic colloids, which requires some local symmetry breaking.…

Soft Condensed Matter · Physics 2021-09-08 Jeanne Decayeux , Vincent Dahirel , Marie Jardat , Pierre Illien

Active colloidal systems with non-equilibrium self-organization is a long-standing, challenging area in biology. To understand how hydrodynamic flow may be used to actively control self-assembly of Janus particles (JPs), we use a model…

Soft Condensed Matter · Physics 2022-08-09 Szu-Pei Fu , Rolf Ryham , Bryan Quaife , Y. -N. Young

Collective behaviors such as swarming, chemical signaling, and clustering are fundamental to biological microorganisms, enabling hierarchical colony formation, coordinated motion, and enhanced nutrient accessibility crucial for their…

Soft Condensed Matter · Physics 2025-02-12 Sobiya Ashraf , Pawan Kumar , Prateek Dwivedi , Frédéric Blanc , Dipin Pillai , Rahul Mangal

A mechanism of formation of small-scale inhomogeneities in spatial distributions of aerosols and droplets associated with clustering instability in the atmospheric turbulent flow is discussed. The particle clustering is a consequence of a…

Atmospheric and Oceanic Physics · Physics 2012-07-26 T. Elperin , N. Kleeorin , M. A. Liberman , V. L'vov , I. Rogachevskii

We study the behaviour of catalytically active droplets in multi-component conserved mixtures affected by noise. Working in the thin interface limit, we analytically determine the state diagram of the system, characterized by multiple…

Soft Condensed Matter · Physics 2025-12-01 Jacopo Romano , Martin Kjøllesdal Johnsrud , Benoît Mahault , Ramin Golestanian

We develop the hydrodynamic theory of dry, polar ordered, active matter (``flocking") with autochemotaxis; i.e., self-propelled entities moving in the same direction, each emitting a substance which attracts the others (e.g., ants). We find…

Soft Condensed Matter · Physics 2024-01-22 Maxx Miller , John Toner

We present an experimental study on the collective behavior of macroscopic self-propelled particles that are externally excited by light. This property allows testing the system response to the excitation intensity in a very versatile…

Soft Condensed Matter · Physics 2025-09-03 Sára Lévay , Axel Katona , Hartmut Löwen , Raúl Cruz Hidalgo , Iker Zuriguel

In a stationary ergodic process, clustering is defined as the tendency of events to appear in series of increased frequency separated by longer breaks. Such behavior, contradicting the theoretical "unbiased behavior" with exponential…

Probability · Mathematics 2008-10-27 Tomasz Downarowicz , Yves Lacroix , Didier Léandri

Collectives of actively-moving particles can spontaneously segregate into dilute and dense phases through a process known as motility-induced phase separation (MIPS). This captivating phenomenon is well-studied for randomly-moving particles…

Soft Condensed Matter · Physics 2026-03-04 Hongbo Zhao , Qiwei Yu , Andrej Košmrlj , Sujit S. Datta

We consider a coarse-grained description of a system of self-propelled particles given by hydrodynamic equations for the density and polarization fields. We find that the ordered moving or flocking state of the system is unstable to spatial…

Statistical Mechanics · Physics 2013-05-29 Shradha Mishra , Aparna Baskaran , M. Cristina Marchetti

The collective chemotaxis of multicellular clusters is an important phenomenon in various physiological contexts, ranging from embryonic development to cancer metastasis. Such clusters often display interesting shape dynamics and…

Tissues and Organs · Quantitative Biology 2025-06-03 Monika Sanoria , Gema Malet-Engra , Giorgio Scita , Nir Gov , Ajay Gopinathan

Phoretic mechanisms, whereby gradients of chemical solutes induce surface-driven flows, have recently been used to generate directed propulsion of patterned colloidal particles. When the chemical solutes diffuse slowly, an instability…

Fluid Dynamics · Physics 2019-12-18 Sébastien Michelin , Simon Game , Eric Lauga , Eric Keaveny , Demetrios Papageourgiou

Self-propelled colloids constitute an important class of intrinsically non-equilibrium matter. Typically, such a particle moves ballistically at short times, but eventually changes its orientation, and displays random-walk behavior in the…

Soft Condensed Matter · Physics 2019-05-27 Jochen Arlt , Vincent A Martinez , Angela Dawson , Teuta Pilizota , Wilson C K Poon

This Article presents a nonequilibrium thermodynamic theory for the mean-field precipitation, aggregation and pattern formation of colloidal clusters. A variable gradient energy coefficient and the arrest of particle diffusion upon…

Soft Condensed Matter · Physics 2018-10-17 Thomas Petersen , Martin Z. Bazant , Roland J. M. Pellenq , Franz-Josef Ulm

A simple three-dimensional model of a fluid whose constituent particles interact via a short range attractive and long range repulsive potential is used to model the aggregation into large spherical-like clusters made up of hundreds of…

Soft Condensed Matter · Physics 2025-02-11 Antonio Díaz-Pozuelo , Diego González-Salgado , Enrique Lomba

After rainfall, pine needles often float on the surface of small puddles. As the water evaporates, they self-assemble into distinct clusters. Motivated by this natural phenomenon, we experimentally investigate the dynamic evolution of…

Soft Condensed Matter · Physics 2026-01-06 Xin Li , Shuchen Zhang , Mark J. Bowick , Duanduan Wan

The dynamics and pair trajectory of two self-propelled colloids are reported. The autonomous motions of the colloids are due to a catalytic chemical reaction taking place asymmetrically on their surfaces that generates a concentration…

Soft Condensed Matter · Physics 2016-06-29 Nima Sharifi-Mood , Ali Mozaffari , Ubaldo Córdova-Figueroa

One standard approach to describe the collective behaviour of self-propelled particles is the Vicsek model: point-like self-propelled particles tend to align their migration directions to the ones of their nearer neighbours at each…

Soft Condensed Matter · Physics 2013-11-27 Andreas M. Menzel

A common feature of biological self-organization is how active agents communicate with each other or their environment via chemical signaling. Such communications, mediated by self-generated chemical gradients, have consequences for both…

Soft Condensed Matter · Physics 2023-11-07 Babak Vajdi Hokmabad , Suropriya Saha , Jaime Agudo-Canalejo , Ramin Golestanian , Corinna C. Maass