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Many active biological particles, such as swimming microorganisms or motor-proteins, do work on their environment by going though a periodic sequence of shapes. Interactions between particles can lead to the phase-synchronization of their…

Soft Condensed Matter · Physics 2022-07-26 Brato Chakrabarti , Michael J. Shelley , Sebastian Fürthauer

Systems of motile microorganisms exhibit a multitude of collective phenomena, including motility-induced phase separation and turbulence. Sensing of the environment and adaptation of movement plays an essential role in the emergent…

Soft Condensed Matter · Physics 2024-06-26 Segun Goh , Elmar Westphal , Roland G. Winkler , Gerhard Gompper

The complex formations exhibited by schooling fish have long been the object of fascination for biologists and physicists. However, the physical and sensory mechanisms leading to organized collective behavior remain elusive. On the physical…

Fluid Dynamics · Physics 2025-07-09 Anand U. Oza , Eva Kanso , Michael J. Shelley

Microswimmers are active particles of microscopic size that self-propel by setting the surrounding fluid into motion. According to the kind of far-field fluid flow that they induce, they are classified into pushers and pullers. Many studies…

Soft Condensed Matter · Physics 2019-08-01 Giorgio Pessot , Hartmut Löwen , Andreas M. Menzel

Three-dimensional simulations with fully resolved hydrodynamics are performed to study the collective motion of model swimmers in confinement. We show that certain swimming mechanisms can lead to traveling wave-like collective motion even…

Soft Condensed Matter · Physics 2016-04-20 Norihiro Oyama , John Jairo Molina , Ryoichi Yamamoto

Populations of swimming microorganisms produce fluid motions that lead to dramatically enhanced diffusion of tracer particles. Using simulations of suspensions of swimming particles in a periodic domain, we capture this effect and show that…

Soft Condensed Matter · Physics 2008-07-22 Patrick T. Underhill , Juan P. Hernández-Ortiz , Michael D. Graham

We investigate the hydrodynamic stability and transport of magnetic microswimmers in an external field using a kinetic theory framework. Combining linear stability analysis and nonlinear 3D continuum simulations, we show that for…

Soft Condensed Matter · Physics 2019-02-20 Fabian R. Koessel , Sara Jabbari-Farouji

Suspensions of swimming micro-organisms are known to undergo intricate collective dynamics as a result of hydrodynamic and collision interactions. Micro-swimmers, such as bacteria and micro-algae, naturally live and have evolved in complex…

Fluid Dynamics · Physics 2025-10-31 Yasser Almoteri , Enkeleida Lushi

To explore and react to their environment, living micro-swimmers have developed sophisticated strategies for locomotion - in particular, motility with multiple gaits. To understand the physical principles associated with such a behavioural…

Some microorganisms, such as spermatozoa, synchronize their flagella when swimming in close proximity. Using a simplified model (two infinite, parallel, two-dimensional waving sheets), we show that phase-locking arises from hydrodynamics…

Biological Physics · Physics 2009-09-15 Gwynn J. Elfring , Eric Lauga

Hydrodynamic interactions (HIs) are important in biophysics research because they influence both the collective and the individual behaviour of microorganisms and self-propelled particles. For instance, HIs at the micro-swimmer level…

Fluid Dynamics · Physics 2022-03-30 Maria Tătulea-Codrean , Eric Lauga

Previous particle-based computer simulations have revealed a significantly more pronounced tendency of spontaneous global polar ordering in puller (contractile) microswimmer suspensions than in pusher (extensile) suspensions. We here…

Soft Condensed Matter · Physics 2018-10-23 Christian Hoell , Hartmut Löwen , Andreas M. Menzel

Colloidal particles moving in a fluid interact via the induced velocity field. The collective dynamic state for a class of actively forced colloids, driven by harmonic potentials via a rule that couples forces to configurations, to perform…

Soft Condensed Matter · Physics 2011-10-11 Loïc Damet , Giovanni M. Cicuta , Jurij Kotar , Marco Cosentino Lagomarsino , Pietro Cicuta

We explore the mechanism underlying the anomalous rheology of puller-type microswimmer suspensions through direct hydrodynamic simulations. Puller-type swimmers generate contractile flow fields along their swimming direction, leading to…

Soft Condensed Matter · Physics 2025-03-04 Haruki Hayano , Akira Furukawa

Geometric confinements are frequently encountered in soft matter systems and in particular significantly alter the dynamics of swimming microorganisms in viscous media. Surface-related effects on the motility of microswimmers can lead to…

The nature of emergent collective behaviors of moving physical agents interacting with their neighborhood is a long-standing open issue in physical and biological systems alike. This calls for studies on the control of synchronization and…

Statistical Mechanics · Physics 2017-08-11 Tirthankar Banerjee , Abhik Basu

Recent experimental studies have shown that confinement can profoundly affect self-organization in semi-dilute active suspensions, leading to striking features such as the formation of steady and spontaneous vortices in circular domains and…

Fluid Dynamics · Physics 2016-08-25 Maxime Theillard , Roberto Alonso-Matilla , David Saintillan

We study the dynamics and interaction of two swimming bacteria, modeled by self-propelled dumbbell-type structures. We focus on alignment dynamics of a coplanar pair of elongated swimmers, which propel themselves either by pushing" or…

Analysis of PDEs · Mathematics 2008-11-24 Vitaliy Gyrya , Igor S. Aranson , Leonid V. Berlyand , Dmitry Karpeev

Low Reynolds number direct simulations of large populations of hydrodynamically interacting swimming particles confined between planar walls are performed. The results of simulations are compared with a theory that describes dilute…

Fluid Dynamics · Physics 2015-05-13 Juan P. Hernandez-Ortiz , Patrick T. Underhill , Michael D. Graham

Active diffusiophoresis - swimming through interaction with a self-generated, neutral, solute gradient - is a paradigm for autonomous motion at the micrometer scale. We study this propulsion mechanism within a linear response theory.…

Statistical Mechanics · Physics 2015-05-30 Benedikt Sabass , Udo Seifert