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We theoretically describe the dynamics of swimmer populations confined in thin liquid films. We first demonstrate that hydrodynamic interactions between confined swimmers only depend on their shape and are independent of their specific…

Soft Condensed Matter · Physics 2015-06-11 Tommaso Brotto , Jean-Baptiste Caussin , Eric Lauga , Denis Bartolo

The complex formations exhibited by schooling fish have long been the object of fascination for biologists and physicists. However, the physical and sensory mechanisms leading to organized collective behavior remain elusive. On the physical…

Fluid Dynamics · Physics 2025-07-09 Anand U. Oza , Eva Kanso , Michael J. Shelley

We derive from first principles a three-dimensional theory of self-propelled particle swarming in a viscous fluid environment. Our model predicts emergent collective behavior that depends critically on fluid opacity, mechanism of…

Soft Condensed Matter · Physics 2026-05-12 Yao-Li Chuang , M. R. D'Orsogna , T. Chou

Self-propelled micron-size particles suspended in a fluid, like bacteria or synthetic microswimmers, are strongly non-equilibrium systems where particle motility breaks the microscopic detailed balance, often resulting in large-scale…

Soft Condensed Matter · Physics 2024-03-15 Dóra Bárdfalvy , Viktor Škultéty , Cesare Nardini , Alexander Morozov , Joakim Stenhammar

We investigate the dynamics of model microswimmers under confinement, in cylindrical geometries, by means of three dimensional direct numerical calculations with fully resolved hydrodynamics. Such swimmers are known to show collective…

Soft Condensed Matter · Physics 2017-09-26 Norihiro Oyama , John Jairo Molina , Ryoichi Yamamoto

Microorganisms living in microfluidic environments often form multi-species swarms, where they can leverage collective motions to achieve enhanced transport and spreading. Nevertheless, there is a general lack of physical understandings of…

Fluid Dynamics · Physics 2022-02-14 Bryce Palmer , Wen Yan , Tong Gao

Fish schooling is often modeled with self-propelled particles subject to phenomenological behavioral rules. Although fish are known to sense and exploit flow features, these models usually neglect hydrodynamics. Here, we propose a novel…

Biological Physics · Physics 2018-05-16 Audrey Filella , François Nadal , Clément Sire , Eva Kanso , Christophe Eloy

The hydrodynamic interactions of a suspension of self-propelled particles are studied using a direct numerical simulation method which simultaneously solves for the host fluid and the swimming particles. A modified version of the "Smoothed…

Soft Condensed Matter · Physics 2013-01-15 John J. Molina , Yasuya Nakayama , Ryoichi Yamamoto

The acoustofluidic method holds great promise for manipulating microorganisms. When exposed to the steady vortex structures of acoustic streaming flow, these microorganisms exhibit intriguing dynamic behaviors, such as hydrodynamic trapping…

Fluid Dynamics · Physics 2025-04-25 Xuyang Sun , Wenchang Tan , Yi Man

Under partial confinement, the motion of colloidal particles is restricted to a plane but their dynamics is influenced by hydrodynamic interactions mediated by the unconfined, three--dimensional flow of the embedding fluid. We demonstrate…

Soft Condensed Matter · Physics 2013-05-17 J. Bleibel , A. Dominguez , F. Günther , J. Harting , M. Oettel

We study the collective motion of confined spherical microswimmers such as active colloids which we model by so-called squirmers. To simulate hydrodynamic flow fields including thermal noise, we use the method of multi-particle collision…

Soft Condensed Matter · Physics 2014-03-20 Andreas Zöttl , Holger Stark

We investigate the effect of cooperative interactions in an ensemble of microorganisms, modelled as self-propelled disk-like and rod-like particles, in a three-dimensional turbulent flow to show flocking as an emergent phenomenon. Building…

Soft Condensed Matter · Physics 2020-05-20 Anupam Gupta , Amal Roy , Arnab Saha , Samriddhi Sankar Ray

Both natural and artificial small-scale swimmers may often self-propel in environments subject to complex geometrical constraints. While most past theoretical work on low-Reynolds number locomotion addressed idealised geometrical…

Fluid Dynamics · Physics 2017-11-16 Alexander Chamolly , Takuji Ishikawa , Eric Lauga

Self-propelled particles with hydrodynamic interactions (microswimmers) have previously been shown to produce long-range ordering phenomena. Many theoretical explanations for these collective phenomena are connected to instabilities in the…

Soft Condensed Matter · Physics 2017-05-03 Yuzhou Qian , Peter R. Kramer , Patrick T. Underhill

Micron-sized self-propelled (active) particles can be considered as model systems for characterizing more complex biological organisms like swimming bacteria or motile cells. We produce asymmetric microswimmers by soft lithography and study…

We consider an Individual-Based Model for self-rotating particles interacting through local alignment and investigate its macroscopic limit. This model describes self-propelled particles moving in the plane and trying to synchronize their…

Mathematical Physics · Physics 2013-06-17 Pierre Degond , Giacomo Dimarco , Thi Bich Ngoc Mac

Suspensions of self-propelled particles, such as swimming micro-organisms, are known to undergo complex dynamics as a result of hydrodynamic interactions. This fluid dynamics video presents a numerical simulation of such a suspension, based…

Fluid Dynamics · Physics 2009-10-20 David Saintillan , Amir Alizadeh Pahlavan

Self-propelled microparticles create flow fields that determine how they interact with surfaces, external flows, and each other. These flow fields fall into distinct classes--pushers, pullers, and neutral swimmers--each exhibiting…

Soft Condensed Matter · Physics 2025-12-25 Lisa Rohde , Gordei Anchutkin , Viktor Holubec , Frank Cichos

We investigate theoretically the collective dynamics of a suspension of low Reynolds number swimmers that are confined to two dimensions by a thin fluid film. Our model swimmer is characterized by internal degrees of freedom which locally…

Biological Physics · Physics 2015-05-19 M. Leoni , T. B. Liverpool

Hydrodynamic interactions between fluid-dispersed particles are ubiquitous in soft matter and biological systems and they give rise to intriguing collective phenomena. While it was reported that these interactions can facilitate…

Soft Condensed Matter · Physics 2022-08-29 Eric Cereceda-López , Dominik Lips , Antonio Ortiz-Ambriz , Artem Ryabov , Philipp Maass , Pietro Tierno
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