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Movement within eukaryotic cells largely originates from localized forces exerted by myosin motors on scaffolds of actin filaments. Although individual motors locally exert both contractile and extensile forces, large actomyosin structures…

Biological Physics · Physics 2014-10-06 Martin Lenz

We investigate the growth of a branched actin network under load. Using a combination of simulations and theory, we show that the network adapts to the load and exhibits two regimes: a finite velocity at low stress, followed by a power-law…

Soft Condensed Matter · Physics 2025-01-10 Magid Badaoui , Serge Dmitrieff

Filopodia are bundles of actin filaments that extend out ahead of the leading edge of a crawling cell to probe its upcoming environment. {\it In vitro} experiments [D. Vignjevic {\it et al.}, J. Cell Biol. {\bf 160}, 951 (2003)] have…

Subcellular Processes · Quantitative Biology 2010-05-14 K. -C. Lee , A. Gopinathan , J. M. Schwarz

We examine the nonequilibrium production of topological defects -- braids -- in semiflexible filament bundles under cycles of compression and tension. During these cycles, the period of compression facilitates the thermally activated pair…

Soft Condensed Matter · Physics 2021-05-28 Valentin M. Slepukhin , Alex J. Levine

The inert polymer polyethylene glycol (PEG) induces a "bundling" phenomenon in F-actin solutions when its concentration exceeds a critical onset value C_o. Over a limited range of PEG molecular weight and ionic strength, C_o can be…

Soft Condensed Matter · Physics 2009-11-10 M. Hosek , J. X. Tang

Using molecular dynamics simulations, we investigate the aggregation behavior of neutral stiff (rod-like) and flexible polymer chains mediated by attractive crowders. Attractive crowders serve as bridging agents, inducing aggregation…

Soft Condensed Matter · Physics 2025-04-02 Hitesh Garg , Satyavani Vemparala

We present a statistical mechanical study of stiff polymers, motivated by experiments on actin filaments and the considerable current interest in polymer networks. We obtain simple, approximate analytical forms for the force-extension…

Soft Condensed Matter · Physics 2009-11-13 Abhijit Ghosh , Joseph Samuel , Supurna Sinha

The actomyosin cytoskeleton, a protein assembly comprising actin fibers and the myosin molecular motor, drives various cellular dynamics through contractile force generation at high densities. However, the relationship between the density…

Soft Condensed Matter · Physics 2025-01-07 Tomoka Kashiwabara , Tatsuya Fukuyama , Yusuke T. Maeda

Current understanding of how contractility emerges in disordered actomyosin networks of non-muscle cells is still largely based on the intuition derived from earlier works on muscle contractility. This view, however, largely overlooks the…

Biological Physics · Physics 2018-04-11 James E. Komianos , Garegin A. Papoian

Cells and tissues exert forces and can actively change shape. This strikingly autonomous behavior is powered by the cytoskeleton, which includes an active gel of actin filaments, crosslinks, and myosin molecular motors. Although individual…

Soft Condensed Matter · Physics 2017-07-26 José Alvarado , Misha Sheinman , Abhinav Sharma , Fred C. MacKintosh , Gijsje H. Koenderink

Amontons' law postulates a monotonic relationship between frictional force and the normal load applied to a sliding contact. This empirical rule, however, fails in systems where internal degrees of freedom - such as structural or electronic…

Materials Science · Physics 2026-03-19 Hongri Gu , Anton Lüders , Clemens Bechinger

A phenomenological description for active bundles of polar filaments is presented. The activity of the bundle results from crosslinks, that induce relative displacements between the aligned filaments. Our generic description is based on…

Statistical Mechanics · Physics 2009-11-07 Karsten Kruse , Frank Julicher

We present a model for the actin contractile ring of adherent animal cells. The model suggests that the actin concentration within the ring and consequently the power that the ring exerts both increase during contraction. We demonstrate the…

Biological Physics · Physics 2010-08-06 D. Biron , E. Alvarez-Lacalle , Tsvi Tlusty , Elisha Moses

The motility of certain gram-negative bacteria is mediated by retraction of type IV pili surface filaments, which are essential for infectivity. The retraction is powered by a strong molecular motor protein, PilT, producing very high forces…

Subcellular Processes · Quantitative Biology 2007-05-23 Martin Linden , Tomi Tuohimaa , Ann-Beth Jonsson , Mats Wallin

A microtubule (MT) is a hollow tube of approximately 25 nm diameter. The two ends of the tube are dissimilar and are designated as `plus' and `minus' ends. Motivated by the collective push and pull exerted by a bundle of MTs during…

Subcellular Processes · Quantitative Biology 2015-01-12 Dipanwita Ghanti , Debashish Chowdhury

The growth of actin filament networks is a fundamental biological process that drives a variety of cellular and intracellular motions. During motility, eukaryotic cells and intracellular pathogens are propelled by actin networks organized…

Biological Physics · Physics 2009-11-13 Joshua W. Shaevitz , Daniel A. Fletcher

We investigate a two-dimensional system of active particles confined to a narrow annular domain. Despite the absence of explicit interactions among the velocities or the active forces of different particles, the system displays a transition…

Statistical Mechanics · Physics 2022-03-15 Lorenzo Caprini , Claudio Maggi , Umberto Marini Bettolo Marconi

Biochemistry and mechanics are closely coupled in cell adhesion. At sites of cell-matrix adhesion, mechanical force triggers signaling through the Rho-pathway, which leads to structural reinforcement and increased contractility in the actin…

Subcellular Processes · Quantitative Biology 2010-02-24 A. Besser , U. S. Schwarz

Myosin II isoforms with varying mechanochemistry and filament size interact with filamentous actin (F-actin) networks to generate contractile forces in cells. How their properties control force production in environments with varying…

Subcellular Processes · Quantitative Biology 2014-07-09 Samantha Stam , Jon Alberts , Margaret L. Gardel , Edwin Munro

We introduce a phenomenological model to study the properties of bundles of polar filaments which interact via active elements. The stability of the homogeneous state, the attractors of the dynamics in the unstable regime and the tensile…

Statistical Mechanics · Physics 2009-10-31 Karsten Kruse , Frank Julicher