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The encounter of diffusing entities underlies a wide range of natural phenomena. The dynamics of these first-passage dynamics are strongly influenced by confining geometries. Confinement modifies microscopic diffusion through conservative…

Soft Condensed Matter · Physics 2026-04-22 Guirec de Tournemire , Nicolas Fares , Yacine Amarouchene , Thomas Salez

Many microorganisms and artificial microswimmers use helical appendages in order to generate locomotion. Though often rotated so as to produce thrust, some species of bacteria such Spiroplasma, Rhodobacter sphaeroides and Spirochetes induce…

Biological Physics · Physics 2018-10-23 Lyndon Koens , Hang Zhang , Martin Moeller , Ahmed Mourran , Eric Lauga

The actomyosin cortex, a thin layer of a cross-linked polymer network near the cell surface, generates active forces that are responsible for cell shape changes. Many developmental processes that involve such cell shape changes, most…

Soft Condensed Matter · Physics 2026-02-26 Da Gao , Alexander Mietke , Rui Ma

Biological microswimmers are known to navigate upstream of an external flow (positive rheotaxis) in trajectories ranging from linear, spiral to oscillatory. Such rheotaxis stems from the interplay between the motion and complex shapes of…

Soft Condensed Matter · Physics 2022-05-27 Ranabir Dey , Carola M. Buness , Babak Vajdi Hokmabad , Chenyu Jin , Corinna C. Maass

Many small organisms self-propel in viscous fluids using travelling wave-like deformation of their bodies or appendages. Examples include small nematodes moving through soil using whole-body undulations or spermatozoa swimming through mucus…

Biological Physics · Physics 2015-07-02 Emily E. Riley , Eric Lauga

The habitat of microorganisms is typically complex and viscoelastic. A natural question arises: Do polymers in a suspension of active swimmers enhance chaotic motion or promote orientational order? We address this issue by performing…

Soft Condensed Matter · Physics 2025-12-23 Yuan Zhou , Qingzhi Zou , Ignacio Pagonabarraga , Kaihuan Zhang , Kai Qi

Confined suspensions of active particles show peculiar dynamics characterized by wall accumulation, as well as upstream swimming, centerline depletion and shear-trapping when a pressure-driven flow is imposed. We use theory and numerical…

Fluid Dynamics · Physics 2023-07-19 Barath Ezhilan , David Saintillan

The motion of microorganisms in their natural habitat is strongly influenced by their propulsion mechanisms, geometrical constraints, and random fluctuations. Here, we study numerically the first-passage-time (FPT) statistics of…

Soft Condensed Matter · Physics 2025-09-19 Yanis Baouche , Magali Le Goff , Thomas Franosch , Christina Kurzthaler

Many self-propelled microorganisms are attracted to surfaces. This makes their dynamics in restricted geometries very different from that observed in the bulk. Swimming along walls is beneficial for directing and sorting cells, but may be…

The locomotion of microorganisms and spermatozoa in complex viscoelastic fluids is of critical importance in many biological processes such as fertilization, infection, and biofilm formation. Depending on their propulsion mechanisms,…

Soft Condensed Matter · Physics 2021-09-14 Gaojin Li , Eric Lauga , Arezoo M. Ardekani

A collection of microswimmers immersed in an incompressible fluid is characterised by strong interactions due to the long-range nature of the hydrodynamic fields generated by individual organisms. As a result, suspensions of rear-actuated…

Soft Condensed Matter · Physics 2023-12-19 Viktor Škultéty , Dóra Bárdfalvy , Joakim Stenhammar , Cesare Nardini , Alexander Morozov

A simple model for the motion of shape-changing swimmers in Poiseuille flow was recently proposed and numerically explored by Omori et al. (2022). These explorations hinted that a small number of interacting mechanics can drive long-time…

We study the motion of a microscopic swimmer composed of a semiflexible polymer anchored at the surface of a magnetic sphere using hydrodynamic simulations and scaling arguments. The swimmer is driven by a rotating magnetic field, and…

Soft Condensed Matter · Physics 2007-05-23 A. Alexander-Katz

In this paper, we give formulas for the swimming of simplified two-dimensional bodies in complex fluids using the reciprocal theorem. By way of these formulas we calculate the swimming velocity due to small-amplitude deformations on the…

Fluid Dynamics · Physics 2016-04-28 Gwynn J. Elfring , Gaurav Goyal

There is increasing evidence that mammalian cells not only crawl on substrates but can also swim in fluids. To elucidate the mechanisms of the onset of motility of cells in suspension, a model which couples actin and myosin kinetics to…

Biological Physics · Physics 2019-09-18 Alexander Farutin , Jocelyn Etienne , Chaouqi Misbah , Pierre Recho

We propose and analyze a simple model for the evolution of an immersed, inextensible filament which incorporates linear viscoelastic effects of the surrounding fluid. The model is a closed-form system of equations along the curve only which…

Analysis of PDEs · Mathematics 2024-05-21 Laurel Ohm

Microswimmers typically operate in complex environments. In biological systems, often diverse species are simultaneously present and interact with each other. Here, we derive a (time-dependent) particle-scale statistical description, namely…

Soft Condensed Matter · Physics 2019-08-13 Christian Hoell , Hartmut Löwen , Andreas M. Menzel

We study the fluid drift due to a time-dependent dumbbell model of a microswimmer. The model captures important aspects of real microswimmers such as a time-dependent flagellar motion and a no-slip body. The model consists of a rigid sphere…

Soft Condensed Matter · Physics 2017-02-01 Peter Mueller , Jean-Luc Thiffeault

The role of passive body dynamics on the kinematics of swimming micro-organisms in complex fluids is investigated. Asymptotic analysis of small amplitude motions of a finite-length undulatory swimmer in a Stokes-Oldroyd-B fluid is used to…

Fluid Dynamics · Physics 2017-10-11 Becca Thomases , Robert D. Guy

Movement is a fundamental behaviour of organisms that brings about beneficial encounters with resources and mates, but at the same time exposes the organism to dangerous encounters with predators. The movement patterns adopted by organisms…

Populations and Evolution · Quantitative Biology 2014-03-26 Giuseppe Bianco , Patrizio Mariani , Andre W. Visser , Maria Grazia Mazzocchi , Simone Pigolotti