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The behavior of fluid interfaces far from equilibrium plays central roles in nature and in industry. Active swimmers trapped at interfaces can alter transport at fluid boundaries with far reaching implications. Swimmers can become trapped…

Soft Condensed Matter · Physics 2024-02-08 Jiayi Deng , Mehdi Molaei , Nicholas G. Chisholm , Kathleen J. Stebe

Recent experimental work has shown that eukaryotic cells can swim in a fluid as well as crawl on a substrate. We investigate the swimming behavior of Dictyostelium discoideum amoebae who swim by initiating traveling protrusions at the front…

Cell Behavior · Quantitative Biology 2015-09-14 Qixuan Wang , Hans G. Othmer

One of the essential functions of living organisms is spontaneous migration through the deformation of their body, such as crawling, swimming, and walking. Depending on the size of the object, the efficient migratory mode should be altered…

Soft Condensed Matter · Physics 2024-02-07 H. Ebata , Y. Nishigami , H. Fujiwara , S. Kidoaki , M. Ichikawa

Eukaryotic swimming cells such as spermatozoa, algae or protozoa use flagella or cilia to move in viscous fluids. The motion of their flexible appendages in the surrounding fluid induces propulsive forces that balance with the viscous drag…

Soft Condensed Matter · Physics 2023-10-06 Maciej Lisicki , Marcos F. Velho Rodrigues , Eric Lauga

Microorganisms ofter move in confined, disordered environments, where hydrodynamic couplings can modify their transport behavior. Using extensive finite-element simulations, we investigate the dynamics of microswimmers -- modeled as…

Soft Condensed Matter · Physics 2026-03-24 Mirko Residori , Sebastian Aland , Christina Kurzthaler

Few simulations exist for microswimmers near deformable interfaces. Here, we present numerical simulations of the hydrodynamic flows associated with a single microswimmer embedded in a binary fluid mixture. The two fluids demix, separated…

Soft Condensed Matter · Physics 2022-05-24 Chao Feng , John J. Molina , Matthew S. Turner , Ryoichi Yamamoto

Active droplets are artificial microswimmers built from a liquid dispersion by microfluidic tools and showing self-propelled motion. These systems hold particular interest for mimicking biological phenomena, such as some aspects of cell…

Soft Condensed Matter · Physics 2023-06-14 Adriano Tiribocchi , Mihir Durve , Marco Lauricella , Andrea Montessori , Davide Marenduzzo , Sauro Succi

Some euglenids, a family of aquatic unicellular organisms, can develop highly concerted, large-amplitude peristaltic body deformations. This remarkable behaviour has been known for centuries. Yet, its function remains controversial, and is…

Soft Condensed Matter · Physics 2020-05-26 Giovanni Noselli , Alfred Beran , Marino Arroyo , Antonio DeSimone

Marine organisms manipulate their surrounding flow through their swimming dynamics, which affects the transport of their own odor cues. We demonstrate by direct numerical simulations how a group of swimmers, moving at intermediate Reynolds…

Fluid Dynamics · Physics 2026-05-04 Martin James , Francesco Viola , Agnese Seminara

The millimeter-long soil-dwelling nematode {\it C. elegans} propels itself by producing undulations that propagate along its body and turns by assuming highly curved shapes. According to our recent study [PLoS ONE \textbf{7}, e40121 (2012)]…

Fluid Dynamics · Physics 2019-01-18 Alejandro Bilbao , Eligiusz Wajnryb , Siva Vanapalli , Jerzy Blawzdziewicz

Many microorganisms take a chiral path while swimming in an ambient uid. In this paper, we study the combined behavior of two chiral swimmers using the well-known squirmer model taking into account chiral asymmetries. In contrast to the…

Soft Condensed Matter · Physics 2022-02-23 P. S. Burada , R. Maity , F. Jülicher

Many swimming bacteria naturally inhabit confined environments, yet how confinement influences their swimming behaviors remains unclear. Here, we combine experiments, continuum modeling and particle-based simulations to investigate…

Biological Physics · Physics 2025-10-13 Da Wei , Shiyuan Hu , Tangmiao Tang , Yaochen Yang , Fanlong Meng , Yi Peng

Biological and artificial microswimmers often have to propel through a variety of environments, ranging from heterogeneous suspending media to strong geometrical confinement. Under confinement, local flow fields generated by microswimmers,…

Soft Condensed Matter · Physics 2024-06-04 Florian A. Overberg , Gerhard Gompper , Dmitry A. Fedosov

Self-propelled micron-size particles suspended in a fluid, like bacteria or synthetic microswimmers, are strongly non-equilibrium systems where particle motility breaks the microscopic detailed balance, often resulting in large-scale…

Soft Condensed Matter · Physics 2024-03-15 Dóra Bárdfalvy , Viktor Škultéty , Cesare Nardini , Alexander Morozov , Joakim Stenhammar

We explore the mechanism underlying the anomalous rheology of puller-type microswimmer suspensions through direct hydrodynamic simulations. Puller-type swimmers generate contractile flow fields along their swimming direction, leading to…

Soft Condensed Matter · Physics 2025-03-04 Haruki Hayano , Akira Furukawa

Swimming in curved spacetimes is a phenomenon whereby free bodies in curved spacetimes are able to propel themselves by performing cyclic internal motions. When originally proposed, it was further suggested that, in the limit of fast…

General Relativity and Quantum Cosmology · Physics 2025-06-12 Rodrigo Andrade e Silva

Deformable boundaries are omnipresent in the habitats of swimming microorganisms, leading to intricate hydroelastic couplings. Employing a perturbation theory, valid for small deformations, we study the swimming dynamics of pushers and…

Soft Condensed Matter · Physics 2025-02-05 Sagnik Garai , Ursy Makanga , Akhil Varma , Christina Kurzthaler

We address the problem of controlling a dynamical system governing the motion of a 3D weighted shape changing body swimming in a perfect fluid. The rigid displacement of the swimmer results from the exchange of momentum between prescribed…

Optimization and Control · Mathematics 2011-03-30 Thomas Chambrion , Alexandre Munnier

Eukaryote cells have a flexible shape, which dynamically changes according to the function performed by the cell. One mechanism for deforming the cell membrane into the desired shape is through the expression of curved membrane proteins.…

Biological Physics · Physics 2018-12-05 Miha Fošnarič , Samo Penič , Aleš Iglič , Veronika Kralj-Iglič , Mitja Drab , Nir Gov

Low Reynolds number swimmers frequently move near boundaries, such as spirochetes moving through porous tissues and sperm navigating the reproductive tract. Furthermore, these microorganisms must often navigate non-Newtonian fluids such as…

Fluid Dynamics · Physics 2023-11-10 D. Gagnon , B. Thomases , R. D. Guy , P. E. Arratia