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Swimming at low Reynolds number in Newtonian fluids is only possible through non-reciprocal body deformations due to the kinematic reversibility of the Stokes equations. We consider here a model swimmer consisting of two linked spheres,…

Fluid Dynamics · Physics 2017-04-26 Babak Nasouri , Aditi Khot , Gwynn J. Elfring

The paper is about the parking 3-sphere swimmer ($\text{sPr}_3$). This is a low-Reynolds number model swimmer composed of three balls of equal radii. The three balls can move along three horizontal axes (supported in the same plane) that…

Optimization and Control · Mathematics 2016-10-18 François Alouges , Giovanni Di Fratta

Trajectory tracking for microswimmers remains a key challenge in microrobotics, where low-Reynolds-number dynamics make control design particularly complex. In this work, we formulate the trajectory tracking problem as an optimal control…

Robotics · Computer Science 2026-02-11 Lucas Palazzolo , Mickaël Binois , Laëtitia Giraldi

We introduce and investigate the wellposedness of a model describing the self-propelled motion of a small abstract swimmer in the 3-D incompressible fluid governed by the nonstationary Stokes equation, typically associated with the low…

Analysis of PDEs · Mathematics 2012-05-08 A. Y. Khapalov

This work aims at finding optimal navigation policies for thin, deformable microswimmers that progress in a viscous fluid by propagating a sinusoidal undulation along their slender body. These active filaments are embedded in a prescribed,…

Fluid Dynamics · Physics 2023-02-13 Zakarya El Khiyati , Raphael Chesneaux , Laetitia Giraldi , Jeremie Bec

The mechanism of swimming at very low Reynolds number conditions is a topic of interest to biologists and engineering community. We develop a novel kinematic model of a slender flexible swimmer which locomotes in a low Reynolds number…

Systems and Control · Computer Science 2018-03-26 Sudin Kadam , Ravi N. Banavar

Many motile biological cells navigate along concentration gradients of signaling molecules: This chemotaxis guides for instance sperm cells from marine invertebrates, which have to find egg cells in the ocean. While chemotaxis has been…

Biological Physics · Physics 2022-09-21 Steffen Lange , Benjamin M. Friedrich

Cilia and flagella are actively bending slender organelles, performing functions such as motility, feeding and embryonic symmetry breaking. We review the mechanics of viscous-dominated microscale flow, including time-reversal symmetry, drag…

Quantitative Methods · Quantitative Biology 2013-09-06 Thomas D. Montenegro-Johnson , Andrew A. Smith , David J. Smith , Daniel Loghin , John R. Blake

In a fluid environment, flagellated microswimmers propel themselves by rotating their flagella. The morphology of these flagella significantly influences forward speed, swimming efficiency, and directional stability, which are critical for…

Fluid Dynamics · Physics 2025-06-25 Baopi Liu , Lu Chen , Wenjun Xu

Metachronal paddling is a swimming strategy in which an organism oscillates sets of adjacent limbs with a constant phase lag, propagating a metachronal wave through its limbs and propelling it forward. This limb coordination strategy is…

Fluid Dynamics · Physics 2025-07-28 Alana A. Bailey , Robert D. Guy

Sperm modulate their flagellar symmetry to navigate through complex physico-chemical environments and achieve reproductive function. Yet it remains elusive how sperm swim forwards despite the inherent asymmetry of several components that…

Biological Physics · Physics 2023-07-21 Xiaomeng Ren , Hermes Bloomfield-Gadêlha

The role of passive body dynamics on the kinematics of swimming micro-organisms in complex fluids is investigated. Asymptotic analysis of small amplitude motions of a finite-length undulatory swimmer in a Stokes-Oldroyd-B fluid is used to…

Fluid Dynamics · Physics 2017-10-11 Becca Thomases , Robert D. Guy

The optimal strategy for a microscopic swimmer to migrate across a linear shear flow is discussed. The two cases, in which the swimmer is located at large distance, and in the proximity of a solid wall, are taken into account. It is shown…

Soft Condensed Matter · Physics 2015-05-18 Piero Olla

Swimming by shape changes at low Reynolds number is widely used in biology and understanding how the efficiency of movement depends on the geometric pattern of shape changes is important to understand swimming of microorganisms and in…

Fluid Dynamics · Physics 2015-07-31 Qixuan Wang , Hans G. Othmer

Many small organisms self-propel in viscous fluids using travelling wave-like deformation of their bodies or appendages. Examples include small nematodes moving through soil using whole-body undulations or spermatozoa swimming through mucus…

Biological Physics · Physics 2015-07-02 Emily E. Riley , Eric Lauga

Both natural and artificial small-scale swimmers may often self-propel in environments subject to complex geometrical constraints. While most past theoretical work on low-Reynolds number locomotion addressed idealised geometrical…

Fluid Dynamics · Physics 2017-11-16 Alexander Chamolly , Takuji Ishikawa , Eric Lauga

Computational scientists have investigated swimming performance across a multitude of different systems for decades. Most models depend on numerous model parameters and performance is sensitive to those parameters. In this paper, parameter…

Fluid Dynamics · Physics 2020-11-24 Nicholas A. Battista

Swimming of a sphere in a viscous incompressible fluid is studied on the basis of the Navier-Stokes equations for wave-type distortions of the spherical shape. At sizable values of the dimensionless scale number the mean swimming velocity…

Fluid Dynamics · Physics 2019-02-20 B. U. Felderhof , R. B. Jones

We consider a swimmer consisting of a collinear assembly of three spheres connected by two slender rods. This swimmer can propel itself forward by varying the lengths of the rods in a way that is not invariant under time reversal. Although…

Fluid Dynamics · Physics 2019-07-09 Babak Nasouri , Andrej Vilfan , Ramin Golestanian

Robotic swimmers are currently a subject of extensive research and development for several underwater applications. Clever design and planning must rely on simple theoretical models that account for the swimmer's hydrodynamics in order to…

Fluid Dynamics · Physics 2018-11-21 Evgenia Virozub , Oren Wiezel , Alon Wolf , Yizhar Or