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Topological defects play a central role in the formation and organization of various biological systems. Historically, such nonequilibrium defects have been mainly studied in the context of homogeneous active nematics. Phase-separated…

Soft Condensed Matter · Physics 2023-07-13 Timo Krüger , Ivan Maryshev , Erwin Frey

Phase separation is a fundamental phenomenon that produces spatially heterogeneous patterns in soft matter. In this Lecture Note we show that phase separation in these materials generally belongs to what we call "viscoelastic phase…

Soft Condensed Matter · Physics 2013-07-08 Hajime Tanaka

Phase separation of intrinsically disordered proteins is important for the formation of membraneless organelles, or biomolecular condensates, which play key roles in the regulation of biochemical processes within cells. In this work, we…

Soft Condensed Matter · Physics 2020-03-18 Antonia Statt , Helena Casademunt , Clifford P. Brangwynne , Athanassios Z. Panagiotopoulos

Fluid phase equilibrium depends on the external constraints imposed on a system. In a closed system with fixed volume, depending on the average density, a vapor bubble may be stable, metastable, or unstable, with respect to the homogeneous…

Soft Condensed Matter · Physics 2026-02-24 Frederic Caupin , Alberto Zaragoza , Miguel A. Gonzalez , Chantal Valeriani

From the mitotic spindle up to tissues and biofilms, many biological systems behave as active droplets, which often break symmetry and change shape spontaneously. Here, I show that active nematic droplets can experience a fingering…

Soft Condensed Matter · Physics 2022-06-07 Ricard Alert

Phase separation is the thermodynamic process that explains how droplets form in multicomponent fluids. These droplets can provide controlled compartments to localize chemical reactions, and reactions can also affect the droplets' dynamics.…

Soft Condensed Matter · Physics 2022-09-02 David Zwicker

The study of liquid crystals at equilibrium has led to fundamental insights into the nature of ordered materials, as well as to practical applications such as display technologies. Active nematics are a fundamentally different class of…

Soft Condensed Matter · Physics 2015-08-19 Stephen J. DeCamp , Gabriel S. Redner , Aparna Baskaran , Michael F. Hagan , Zvonimir Dogic

We consider a collection of self-driven apolar particles on a substrate that organize into an active nematic phase at sufficiently high density or low noise. Using the dynamical renormalization group, we systematically study the 2d…

Soft Condensed Matter · Physics 2018-02-07 Suraj Shankar , Sriram Ramaswamy , M. Cristina Marchetti

We use linear stability analysis and hybrid lattice Boltzmann simulations to study the dynamical behaviour of an active nematic confined in a channel made of viscoelastic material. We find that the quiescent, ordered active nematic is…

Soft Condensed Matter · Physics 2023-12-20 Francesco Mori , Saraswat Bhattacharyya , Julia M. Yeomans , Sumesh P. Thampi

We consider a continuum model of active viscoelastic matter, whereby an active nematic liquid-crystal is coupled to a minimal model of polymer dynamics with a viscoelastic relaxation time $\tau_C$. To explore the resulting interplay between…

Soft Condensed Matter · Physics 2016-03-30 E. J. Hemingway , M. E. Cates , S. M. Fielding

Phase separating systems that are maintained away from thermodynamic equilibrium via molecular processes represent a class of active systems, which we call active emulsions. These systems are driven by external energy input for example…

Soft Condensed Matter · Physics 2019-06-07 Christoph A. Weber , David Zwicker , Frank Jülicher , Chiu Fan Lee

We investigate experimentally the dynamic phase transition of a two-dimensional active nematic layer interfaced with a passive liquid crystal. Under a temperature ramp that leads to the transition of the passive liquid into a highly…

The emergence of hydrodynamic bend instabilities in ordered suspensions of active particles is widely observed across diverse living and synthetic systems, and is considered to be governed by dipolar active stresses generated by the…

Soft Condensed Matter · Physics 2026-01-09 Sameer Kumar , Niels de Graaf Sousa , Amin Doostmohammadi

We consider a coarse-grained description of a system of self-propelled particles given by hydrodynamic equations for the density and polarization fields. We find that the ordered moving or flocking state of the system is unstable to spatial…

Statistical Mechanics · Physics 2013-05-29 Shradha Mishra , Aparna Baskaran , M. Cristina Marchetti

Recent studies of the phase diagram for spherical, purely repulsive, active particles established the existence of a transition from a liquid-like to a solid-like phase analogous to the one observed in colloidal systems at thermal…

Statistical Mechanics · Physics 2020-04-30 Lorenzo Caprini , Claudio Maggi , Umberto Marini Bettolo Marconi , Matteo Paoluzzi , Andrea Puglisi

Liquid crystal elastomers realize a fascinating new form of soft matter that is a composite of a conventional crosslinked polymer gel (rubber) and a liquid crystal. These {\em solid} liquid crystal amalgams, quite similarly to their…

Soft Condensed Matter · Physics 2009-09-29 Xiangjun Xing , Leo Radzihovsky

The interplay between active matter and its environment is central to understanding emergent behavior in biological and synthetic systems. Here, we show that coupling active nematic flows to small-amplitude deformations of a compliant…

Soft Condensed Matter · Physics 2026-01-19 Varun Venkatesh , Amin Doostmohammadi

We study phase separation between coexisting active and passive fluids in three-dimensions, using numerical simulation and experiments. Chaotic flows of the active phase drive giant interfacial deformations, causing the co-existing phases…

Soft Condensed Matter · Physics 2026-02-16 Paarth Gulati , Liang Zhao , Michio Tateno , Omar A. Saleh , Zvonimir Dogic , M. Cristina Marchetti

We present in this paper a detailed analysis of the flexoelectric instability of a planar nematic layer in the presence of an alternating electric field (frequency $\omega$), which leads to stripe patterns (flexodomains) in the plane of the…

Soft Condensed Matter · Physics 2015-05-27 Alexei Krekhov , Werner Pesch , Agnes Buka

The spontaneous emergence of collective flows is a generic property of active fluids and often leads to chaotic flow patterns characterised by swirls, jets, and topological disclinations in their orientation field. However, the ability to…

Soft Condensed Matter · Physics 2017-03-07 Tyler N. Shendruk , Amin Doostmohammadi , Kristian Thijssen , Julia M. Yeomans