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The field of active nematics has traditionally employed descriptions based on dipolar activity, with interactions that align along a single axis. However, it has been theoretically predicted that interactions with a substrate, prevalent in…

Active drops emit/absorb chemical solutes, whose concentration gradients cause interfacial flows driving their own transport and the propulsion of the droplet. Such non-linear coupling enables active drops to achieve directed…

Fluid Dynamics · Physics 2022-12-01 Nikhil Desai , Sebastien Michelin

Cell motility in viscous fluids is ubiquitous and affects many biological processes, including reproduction, infection, and the marine life ecosystem. Here we review the biophysical and mechanical principles of locomotion at the small…

Soft Condensed Matter · Physics 2009-09-16 Eric Lauga , Thomas R. Powers

We report on the emergence of spontaneously rotating clusters in active emulsions. Ensembles of self-propelling droplets sediment and then self-organise into planar, hexagonally ordered clusters which hover over the container bottom while…

Soft Condensed Matter · Physics 2022-04-25 Babak Vajdi Hokmabad , Akinori Nishide , Prashanth Ramesh , Corinna C. Maass

Defect dynamics in a thin active nematic layer is studied by asymptotic matching of solutions in the defect core and the far field. The analysis is facilitated by the correspondence between the 2D nematic and complex scalar field models.…

Soft Condensed Matter · Physics 2015-06-16 L. M. Pismen

This paper considers the interaction between two droplets placed on a substrate in immediate vicinity. We show here that when the two droplets are of different fluids and especially when one of the droplet is highly volatile, a wealth of…

Fluid Dynamics · Physics 2015-06-11 Mathieu Sellier , Volker Nock , Cécile Gaubert , Claude Verdier

The symmetry breaking of the actin network from radial to longitudinal symmetry has been identified as the major mechanism for keratocytes (fish cells) motility on solid substrate. For strong friction coefficient, the two dimensional actin…

Soft Condensed Matter · Physics 2011-02-16 M. Ben Amar , O. V. Manyuhina , G. Napoli

The emergence of long-range spatiotemporal order from intrinsic chaos is a central challenge in far-from-equilibrium physics. In active fluids, such as cytoskeletal networks driving cellular motion, self-generated flows typically produce…

The genome contains genetic information essential for cell's life. The genome's spatial organization inside the cell nucleus is critical for its proper function including gene regulation. The two major genomic compartments -- euchromatin…

Soft Condensed Matter · Physics 2026-04-24 S. Alex Rautu , Alexandra Zidovska , David Saintillan , Michael J. Shelley

When interacting motile units self-organize into flocks, they realize one of the most robust ordered state found in nature. However, after twenty five years of intense research, the very mechanism controlling the ordering dynamics of both…

Soft Condensed Matter · Physics 2021-10-04 Amélie Chardac , Ludwig A. Hoffmann , Yoann Poupart , Luca Giomi , Denis Bartolo

Topological defects in nematically aligned cell populations play a critical role in modulating collective motion, from microbial colonies to epithelial tissues. Despite the potential of manipulating such topological defects to control…

Soft Condensed Matter · Physics 2023-07-25 Ryo Ienaga , Kazusa Beppu , Yusuke T. Maeda

The actin cytoskeleton is remarkably adaptable and multifunctional. It often organizes into nematic bundles such as contractile rings or stress fibers. However, how a uniform and isotropic actin gel self-organizes into dense nematic bundles…

Biological cells can actively tune their intracellular architecture according to their overall shape. Here we explore the rheological implication of such coupling in a minimal model of a dense cellular material where each cell exerts an…

Soft Condensed Matter · Physics 2022-04-13 Shao-Zhen Lin , Matthias Merkel , Jean-François Rupprecht

Active nematic models explain the topological defects and flow patterns observed in epithelial tissues, but the nature of active stress-whether it is extensile or contractile, a key parameter of the theory-is not well established…

Soft Condensed Matter · Physics 2025-11-11 Jan Rozman , Sumesh P. Thampi , Julia M. Yeomans

One of the defining features of active nematics is that above a critical activity the quiescent state becomes unstable to a distorted, flowing one. We show that spatial variations in activity can fundamentally change the nature of this…

Soft Condensed Matter · Physics 2025-12-15 Alexander J. H. Houston , Michael Grinfeld , Geoff McKay , Nigel J. Mottram

Theories of self-organized active fluid surfaces have emerged as an important class of minimal models for the shape dynamics of biological membranes, cells and tissues. However, due to their inherent geometric nonlinearities and the absence…

Soft Condensed Matter · Physics 2025-07-16 Da Gao , Huayang Sun , Rui Ma , Alexander Mietke

We describe a numerical investigation of a continuum model of an active nematic, concentrating on the regime of active turbulence. Results are presented for the effect of three parameters, activity, elastic constant and rotational diffusion…

Soft Condensed Matter · Physics 2015-06-18 Sumesh P. Thampi , Ramin Golestanian , Julia M. Yeomans

Many of the biological phenomena involve collective dynamics driven by self-propelled motion and nonequilibrium force (i.e., activity) that result in features unexpected from equilibrium physics. On the other hand, biological experiments…

Soft Condensed Matter · Physics 2020-09-08 Lisa Yamauchi , Tomoya Hayata , Masahito Uwamichi , Tomoki Ozawa , Kyogo Kawaguchi

Biological tissues transform between solid-like and liquid-like states in many fundamental physiological events. Recent experimental observations further suggest that in two-dimensional epithelial tissues these solid-liquid transformations…

Soft Condensed Matter · Physics 2024-05-06 Yiwen Tang , Siyuan Chen , Mark J. Bowick , Dapeng Bi

We construct the equations of motion for the coupled dynamics of order parameter and concentration for the nematic phase of driven particles on a solid surface, and show that they imply (i) giant number fluctuations, with a standard…

Soft Condensed Matter · Physics 2009-11-07 Sriram Ramaswamy , R. Aditi Simha , John Toner
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