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Active colloids self-organise to a variety of collective states, ranging from highly motile 'molecules' to complex 3D structures. Using large-scale simulations, we show that hydrodynamic interactions, together with a gravity-like aligning…

Soft Condensed Matter · Physics 2020-03-31 Zaiyi Shen , Alois Würger , Juho S. Lintuvuori

We study the universal behavior of a class of active colloids whose design is inspired by the collective dynamics of natural systems like schools of fish and flocks of birds. These colloids, with off-center repulsive interaction sites,…

Soft Condensed Matter · Physics 2024-10-10 Amir Nourhani

We review recent work on active colloids or swimmers, such as self-propelled microorganisms, phoretic colloidal particles, and artificial micro-robotic systems, moving in fluid-like environments. These environments can be water-like and…

Biological Physics · Physics 2016-02-09 Alison E. Patteson , Arvind Gopinath , Paulo E. Arratia

We review recent experimental, numerical, and analytical results on active suspensions of self-propelled colloidal beads moving in (quasi) two dimensions. Active colloids form part of the larger theme of active matter, which is noted for…

Soft Condensed Matter · Physics 2014-09-03 Julian Bialké , Thomas Speck , Hartmut Löwen

Many microorganisms take a chiral path while swimming in an ambient uid. In this paper, we study the combined behavior of two chiral swimmers using the well-known squirmer model taking into account chiral asymmetries. In contrast to the…

Soft Condensed Matter · Physics 2022-02-23 P. S. Burada , R. Maity , F. Jülicher

When swimming animals form cohesive groups, they can reap several benefits. Our understanding of collective animal motion has traditionally been driven by models based on phenomenological behavioral rules, but more recent work has…

Fluid Dynamics · Physics 2024-11-15 Mohamed Niged Mabrouk , Daniel Floryan

Squirmers that are bottom-heavy experience a torque that aligns them along the vertical so that they swim upwards. In a suspension of many squirmers, they also interact hydrodynamically via flow fields that are initiated by their swimming…

Soft Condensed Matter · Physics 2022-01-11 Felix Rühle , Arne W. Zantop , Holger Stark

Microswimmers are often found in heterogeneous and crowded environments within narrow conduits under external flow conditions, enabling them to perform interesting translational and rotational maneuvers, such as swimming in the upstream…

Soft Condensed Matter · Physics 2026-05-26 Margam Ramprasad , Shubhadeep Mandal , Pallab Sinha Mahapatra

Three-dimensional simulations with fully resolved hydrodynamics are performed to study the collective motion of model swimmers in confinement. We show that certain swimming mechanisms can lead to traveling wave-like collective motion even…

Soft Condensed Matter · Physics 2016-04-20 Norihiro Oyama , John Jairo Molina , Ryoichi Yamamoto

From the formation of animal flocks to the emergence of coordinate motion in bacterial swarms, at all scales populations of motile organisms display coherent collective motion. This consistent behavior strongly contrasts with the difference…

Soft Condensed Matter · Physics 2013-11-11 Antoine Bricard , Jean-Baptiste Caussin , Nicolas Desreumaux , Olivier Dauchot , Denis Bartolo

Despite a mounting evidence that the same gradients which active colloids use for swimming, induce important cross-interactions (phoretic interaction), they are still ignored in most many-body descriptions, perhaps to avoid complexity and a…

Soft Condensed Matter · Physics 2019-02-20 B. Liebchen , H. Löwen

Both natural and artificial small-scale swimmers may often self-propel in environments subject to complex geometrical constraints. While most past theoretical work on low-Reynolds number locomotion addressed idealised geometrical…

Fluid Dynamics · Physics 2017-11-16 Alexander Chamolly , Takuji Ishikawa , Eric Lauga

We perform hydrodynamic simulations using the method of multi-particle collision dynamics and a theoretical analysis to study a single squirmer microswimmer at high P\'eclet number, which moves in a low Reynolds number fluid and under…

Fluid Dynamics · Physics 2018-03-14 Felix Rühle , Johannes Blaschke , Jan-Timm Kuhr , Holger Stark

The effect of gravity on the collective motion of living microswimmers, such as bacteria and micro-algae, is pivotal to unravel not only bio-convection patterns but also the settling of bacterial biofilms on solid surfaces. In this work, we…

Soft Condensed Matter · Physics 2025-04-07 C. Miguel Barriuso G. , Horacio Serna , Ignacio Pagonabarraga , Chantal Valeriani

Understanding how microscopic motility shapes emergent collective behaviors is a challenging task in active matter, especially when self-propulsion is regulated by external cues or via quorum-sensing interactions. To address this problem,…

Statistical Mechanics · Physics 2026-04-13 Alberto Dinelli , Pietro Luigi Muzzeddu

In this paper, we explore the hydrodynamics of spheroidal active particles in viscosity gradients. This work provides a more accurate modeling approach, in comparison to spherical particles, for anisotropic organisms like Paramecium…

Fluid Dynamics · Physics 2025-07-09 Jiahao Gong , Vaseem A. Shaik , Gwynn J. Elfring

Locomotion and transport of microorganisms in fluids is an essential aspect of life. Search for food, orientation toward light, spreading of off-spring, and the formation of colonies are only possible due to locomotion. Swimming at the…

Biological Physics · Physics 2015-05-26 Jens Elgeti , Roland G. Winkler , Gerhard Gompper

We report computer simulation results on the cluster formation of dipolar colloidal particles driven by a rotating external field in a quasi-two-dimensional setup. We focus on the interplay between permanent dipolar and hydrodynamic…

Soft Condensed Matter · Physics 2015-06-12 Sebastian Jäger , Holger Stark , Sabine H. L. Klapp

Microorganisms ofter move in confined, disordered environments, where hydrodynamic couplings can modify their transport behavior. Using extensive finite-element simulations, we investigate the dynamics of microswimmers -- modeled as…

Soft Condensed Matter · Physics 2026-03-24 Mirko Residori , Sebastian Aland , Christina Kurzthaler

The complex formations exhibited by schooling fish have long been the object of fascination for biologists and physicists. However, the physical and sensory mechanisms leading to organized collective behavior remain elusive. On the physical…

Fluid Dynamics · Physics 2025-07-09 Anand U. Oza , Eva Kanso , Michael J. Shelley