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Active matter has been intensely studied for its wealth of intriguing properties such as collective motion, motility-induced phase separation (MIPS), and giant fluctuations away from criticality. However, the precise connection of active…

Statistical Mechanics · Physics 2019-04-17 Juliane U. Klamser , Sebastian C. Kapfer , Werner Krauth

From the mitotic spindle up to tissues and biofilms, many biological systems behave as active droplets, which often break symmetry and change shape spontaneously. Here, I show that active nematic droplets can experience a fingering…

Soft Condensed Matter · Physics 2022-06-07 Ricard Alert

Models of active nematics in biological systems normally require complexity arising from the hydrodynamics involved at the microscopic level as well as the viscoelastic nature of the system. Here we show that a minimal, space-independent,…

Soft Condensed Matter · Physics 2022-06-27 Emmanuel L. C. VI M. Plan , Huong Le Thi , Julia M. Yeomans , Amin Doostmohammadi

We numerically investigate the morphology and disclination line dynamics of active nematic droplets in three dimensions. Although our model only incorporates the simplest possible form of achiral active stress, active nematic droplets…

Soft Condensed Matter · Physics 2021-04-07 Liam J. Ruske , Julia M. Yeomans

Active matter systems evade the constraints of thermal equilibrium, leading to the emergence of intriguing collective behavior. A paradigmatic example is given by motor-filament mixtures, where the motion of motor proteins drives alignment…

Soft Condensed Matter · Physics 2024-09-11 Filippo De Luca , Ivan Maryshev , Erwin Frey

The self-propulsion of +1/2 topological defects is a hallmark of active nematic fluids, where the defects are advected by the flow field they themselves generate. In this paper we propose a minimal model for defect self-propulsion in a…

Soft Condensed Matter · Physics 2025-08-08 Fridtjof Brauns , Myles O'Leary , Arthur Hernandez , Mark J. Bowick , M. Cristina Marchetti

The design of artificial microswimmers is often inspired by the strategies of natural microorganisms. Many of these creatures exploit the fact that elasticity breaks the time-reversal symmetry of motion at low Reynolds numbers, but this…

Machines enabled the Industrial Revolution and are central to modern technological progress: A machine's parts transmit forces, motion, and energy to one another in a predetermined manner. Today's engineering frontier, building artificial…

Soft Condensed Matter · Physics 2021-11-18 A. Aubret , Q. Martinet , J. Palacci

We study numerically the behaviour of a mixture of a passive isotropic fluid and an active polar gel, in the presence of a surfactant favouring emulsification. Focussing on parameters for which the underlying free energy favours the…

The cytoskeleton is a model active matter system that controls diverse cellular processes from division to motility. While both active actomyosin dynamics and actin-microtubule interactions are key to the cytoskeleton's versatility and…

A number of micro-scale biological flows are characterized by spatio-temporal chaos. These include dense suspensions of swimming bacteria, microtubule bundles driven by motor proteins, and dividing and migrating confluent layers of cells. A…

Fluid Dynamics · Physics 2017-08-02 Javier Urzay , Amin Doostmohammadi , Julia M. Yeomans

Droplets are essential for spatially controlling biomolecules in cells. To work properly, cells need to control the emergence and morphology of droplets. On the one hand, driven chemical reactions can affect droplets profoundly. For…

Soft Condensed Matter · Physics 2024-03-14 Noah Ziethen , David Zwicker

We experimentally investigate the dynamics of synthetic active particles composed of gravitationally bouncing, superwalking droplets confined within an annular fluid bath. Driven by a topologically pumping dual-frequency waveform, the…

Fluid Dynamics · Physics 2025-12-19 Rudra Sekhri , Rahil N. Valani , Tapio Simula

Melting of two-dimensional (2D) equilibrium crystals, from superconducting vortex lattices to colloidal structures, is a complex phenomenon characterized by the sequential loss of positional and orientational order. Whereas melting…

Soft Condensed Matter · Physics 2021-10-27 Martin James , Dominik Anton Suchla , Jörn Dunkel , Michael Wilczek

At the heart of the structured architecture and complex dynamics of biological systems are specific and timely interactions operated by biomolecules. In many instances, biomolecular agents are spatially confined to flexible lipid membranes…

Soft Condensed Matter · Physics 2020-01-08 Bortolo Mognetti , Pietro Cicuta , Lorenzo Di Michele

Active matter refers to a broad class of non-equilibrium systems where energy is continuously injected at the level of individual ``particles". These systems exhibit emergent collective behaviors that have no direct thermal-equilibrium…

Soft Condensed Matter · Physics 2026-02-10 Zihan Zheng , Cunyuan Jiang , Yangrui Chen , Matteo Baggioli , Jie Zhang

Active systems, from bacterial suspensions to cellular monolayers, are continuously driven out of equilibrium by local injection of energy from their constituent elements and exhibit turbulent-like and chaotic patterns. Here we demonstrate…

Soft Condensed Matter · Physics 2016-02-04 Amin Doostmohammadi , Michael F. Adamer , Sumesh P. Thampi , Julia M. Yeomans

Active matter embraces systems that self-organize at different length and time scales, often exhibiting turbulent flows. Here, we use a quasi-two-dimensional nematically ordered layer of a protein-based active gel to experimentally…

Soft Condensed Matter · Physics 2017-11-01 Pau Guillamat , Jordi Ignés-Mullol , Francesc Sagués

The adaptive and surprising emergent properties of biological materials self-assembled in far-from-equilibrium environments serve as an inspiration for efforts to design nanomaterials and their properties. In particular, controlling the…

Statistical Mechanics · Physics 2023-07-07 Shriram Chennakesavalu , Sreekanth K. Manikandan , Frank Hu , Grant M. Rotskoff

Self-propulsion is a quintessential aspect of biological systems, which can induce nonequilibrium phenomena that have no counterparts in passive systems. Motivated by biophysical interest together with recent advances in experimental…

Soft Condensed Matter · Physics 2026-02-26 Kazuki Sone , Kazuki Yokomizo , Kyogo Kawaguchi , Yuto Ashida
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