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A striking feature of the collective behavior of spherical microswimmers is that for sufficiently strong self-propulsion they phase-separate into a dense cluster coexisting with a low-density dis- ordered surrounding. Extending our previous…

Soft Condensed Matter · Physics 2016-10-31 Johannes Blaschke , Maurice Maurer , Karthik Menon , Andreas Zöttl , Holger Stark

We numerically investigate the motion of active artificial microswimmers diffusing in a fuel concentration gradient. We observe that, in the steady state, their probability density accumulates in the low-concentration regions, whereas a…

Soft Condensed Matter · Physics 2025-08-18 Pulak K. Ghosh , Yunyun Li , Fabio Marchesoni , Franco Nori

Cooperative motion in biological microswimmers is crucial for their survival as it facilitates adhesion to surfaces, formation of hierarchical colonies, efficient motion, and enhanced access to nutrients. Synthetic microswimmers currently…

Soft Condensed Matter · Physics 2022-04-12 Stefania Ketzetzi , Melissa Rinaldin , Pim Dröge , Joost de Graaf , Daniela J. Kraft

Synthetic microswimmers show great promise in biomedical applications such as drug delivery and microsurgery. Their locomotion, however, is subject to stringent constraints due to the dominance of viscous over inertial forces at low…

Fluid Dynamics · Physics 2020-07-15 Alan Cheng Hou Tsang , Pun Wai Tong , Shreyes Nallan , On Shun Pak

We use high-speed x-ray imaging to reveal how a small (~10cm) desert dwelling lizard, the sandfish (Scincus scincus), swims within a granular medium [1]. On the surface, the lizard uses a standard diagonal gait, but once below the surface,…

Fluid Dynamics · Physics 2009-10-20 Ryan D. Maladen , Yang Ding , Adam Kamor , Daniel I. Goldman

A variety of swimming microorganisms, called ciliates, exploit the bending of a large number of small and densely-packed organelles, termed cilia, in order to propel themselves in a viscous fluid. We consider a spherical envelope model for…

Fluid Dynamics · Physics 2011-08-30 Sebastien Michelin , Eric Lauga

Recent advancements in soft actuators have enabled soft continuum swimming robots to achieve higher efficiency and more closely mimic the behaviors of real marine animals. However, optimizing the design and control of these soft continuum…

Robotics · Computer Science 2025-04-29 Yanhao Yang , Ross L. Hatton

We study the dynamics of gyrotactic microswimmers suspended in homogeneous and isotropic turbulence by using direct numerical simulations (DNS). The swimmers are characterized by three non-dimensional parameters: their aspect ratio…

Soft Condensed Matter · Physics 2026-04-24 Suraj Kumar Nayak , Vishwanath Shukla , Akshay Bhatnagar

Populations of swimming microorganisms produce fluid motions that lead to dramatically enhanced diffusion of tracer particles. Using simulations of suspensions of swimming particles in a periodic domain, we capture this effect and show that…

Soft Condensed Matter · Physics 2008-07-22 Patrick T. Underhill , Juan P. Hernández-Ortiz , Michael D. Graham

We employ three numerical methods to explore the motion of low Reynolds number swimmers, modeling the hydrodynamic interactions by means of the Oseen tensor approximation, lattice Boltzmann simulations and multiparticle collision dynamics.…

Soft Condensed Matter · Physics 2007-05-23 David J. Earl , C. M. Pooley , J. F. Ryder , Irene Bredberg , J. M. Yeomans

In Stokes flow, Purcell's scallop theorem forbids objects with time-reversible (reciprocal) swimming strokes from moving. In the presence of inertia, this restriction is eased and reciprocally deforming bodies can swim. A number of recent…

Fluid Dynamics · Physics 2022-11-30 Nicholas J. Derr , Thomas Dombrowski , Chris H. Rycroft , Daphne Klotsa

Many microorganisms swim in a highly heterogeneous environment with obstacles such as fibers or polymers. To better understand how this environment affects microorganism swimming, we study propulsion of a cylinder or filament in a fluid…

Fluid Dynamics · Physics 2016-04-13 Nguyenho Ho , Karin Leiderman , Sarah D. Olson

Microorganisms ofter move in confined, disordered environments, where hydrodynamic couplings can modify their transport behavior. Using extensive finite-element simulations, we investigate the dynamics of microswimmers -- modeled as…

Soft Condensed Matter · Physics 2026-03-24 Mirko Residori , Sebastian Aland , Christina Kurzthaler

Micro-organisms expend energy moving through complex media. While propulsion speed is an important property of locomotion, efficiency is another factor that may determine the swimming gait adopted by a micro-organism in order to locomote in…

Fluid Dynamics · Physics 2017-12-14 Herve Nganguia , Kyle Pietrzyk , On Shun Pak

In this paper we study swimming of a model organism, the so-called Taylor's swimming sheet, in a viscoelastic fluid close to a solid boundary. This situation comprises natural habitats of many swimming microorganisms, and while previous…

Fluid Dynamics · Physics 2018-01-29 Thomas R. Ives , Alexander Morozov

In nature, metachronal coordination is an efficient strategy for fluid pumping and self-propulsion. Yet, mimetic systems for this type of organization are scarce. Recently, metachronal motion was observed in a bead-based magnetocapillary…

Soft Condensed Matter · Physics 2023-11-07 Sebastian Ziegler , Megan Delens , Ylona Collard , Maxime Hubert , Nicolas Vandewalle , Ana-Sunčana Smith

Geometric confinement plays an important role in the dynamics of natural and synthetic microswimmers from bacterial cells to self-propelled particles in high-throughput microfluidic devices. However, little is known about the effects of…

Fluid Dynamics · Physics 2014-11-13 Alan Cheng Hou Tsang , Eva Kanso

Active dumbbell suspensions constitute one of the simplest model system for collective swimming at low Reynolds number. Generalizing recent work, we derive and analyze stroke-averaged equations of motion that capture the effective…

Soft Condensed Matter · Physics 2010-10-12 Victor B. Putz , Jörn Dunkel

In this paper we study the motion of three linked ellipses moving through a viscous fluid in two dimensions. The angles between the ellipses change with time in a specified manner (the gait) and the resulting time varying configuration is…

Fluid Dynamics · Physics 2015-05-14 J. B. Kajtar , J. J. Monaghan

In this communication we address some hydrodynamic aspects of recently revisited drift mechanism of biogenic mixing in the ocean (Katija and Dabiri, Nature vol. 460, pp. 624-626, 2009). The relevance of the locomotion gait at various…

Fluid Dynamics · Physics 2015-05-19 A. M. Leshansky , L. M. Pismen
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