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Collective locomotion of swimming and flying animals is fascinating in terms of individual-level fluid mechanics and group-level structure and dynamics. Here we bridge and relate these scales through a model of formation flight that views…

Fluid Dynamics · Physics 2025-06-18 Christiana Mavroyiakoumou , Jiajie Wu , Leif Ristroph

We undertake a systematic numerical exploration of self-organized states in a deterministic model of interacting self-propelled particles in two dimensions. In the process, we identify various types of collective motion, namely, disordered…

Statistical Mechanics · Physics 2015-03-19 Jihad Touma , Amer Shreim , Leonid Klushin

We present the hydrodynamic theory of coherent collective motion ("flocking") at a solid-liquid interface, and many of its predictions for experiment. We find that such systems are stable, and have long-range orientational order, over a…

Soft Condensed Matter · Physics 2022-01-05 Niladri Sarkar , Abhik Basu , John Toner

We study the large-time behavior of hydrodynamic model which describes the collective behavior of continuum of agents, driven by pairwise alignment interactions with additional external potential forcing. The external force tends to compete…

Analysis of PDEs · Mathematics 2020-07-15 Ruiwen Shu , Eitan Tadmor

Systems of self-propelled particles are known for their tendency to aggregate and to display swarm behavior. We investigate two model systems, self-propelled rods interacting via volume exclusion, and sinusoidally-beating flagella embedded…

Biological Physics · Physics 2010-09-28 Yingzi Yang , Vincent Marceau , Gerhard Gompper

We present molecular dynamics simulations on the slow dynamics of a mixture of big and small soft-spheres with a large size disparity. Dynamics are investigated in a broad range of temperature and mixture composition. As a consequence of…

Soft Condensed Matter · Physics 2009-11-11 Angel J. Moreno , Juan Colmenero

Fish, birds, insects and robots frequently swim or fly in groups. During their 3 dimensional collective motion, these agents do not stop, they avoid collisions by strong short-range repulsion, and achieve group cohesion by weak long-range…

Soft Condensed Matter · Physics 2018-07-04 Illes J. Farkas , Shuo-Hong Wang

Two hallmarks of non-equilibrium systems, from active colloids to animal herds, are agents motility and nonreciprocal interactions. Their interplay creates feedback loops leading to complex spatiotemporal dynamics crucial to understand and…

Statistical Mechanics · Physics 2025-09-17 David Martin , Daniel Seara , Yael Avni , Michel Fruchart , Vincenzo Vitelli

We provide a bird's eye view on developments in analyzing the long time, large crowd behavior of Cucker-Smale alignment dynamics. We consider a class of (fully-)discrete models, paying particular attention to general alignment protocols in…

Dynamical Systems · Mathematics 2023-06-06 Eitan Tadmor

From the formation of animal flocks to the emergence of coordinate motion in bacterial swarms, at all scales populations of motile organisms display coherent collective motion. This consistent behavior strongly contrasts with the difference…

Soft Condensed Matter · Physics 2013-11-11 Antoine Bricard , Jean-Baptiste Caussin , Nicolas Desreumaux , Olivier Dauchot , Denis Bartolo

The aim of the present paper is to elucidate the transition from collective to random behavior exhibited by various mathematical models of bird flocking. In particular, we compare Vicsek's model [Viscek et al., Phys. Rev. Lett. 75, 1226 --…

Adaptation and Self-Organizing Systems · Physics 2015-06-17 H. Christodoulidi , K. van der Weele , Ch. G. Antonopoulos , T. Bountis

A striking feature of the collective behavior of spherical microswimmers is that for sufficiently strong self-propulsion they phase-separate into a dense cluster coexisting with a low-density dis- ordered surrounding. Extending our previous…

Soft Condensed Matter · Physics 2016-10-31 Johannes Blaschke , Maurice Maurer , Karthik Menon , Andreas Zöttl , Holger Stark

The collective dynamics of externally driven $N_{\rm p}$-colloidal systems ($1\le N_{\rm p}\le 4$) in a confined viscous fluid have been investigated using three-dimensional direct numerical simulations with fully resolved hydrodynamics.…

Soft Condensed Matter · Physics 2018-04-04 Norihiro Oyama , Kosuke Teshigawara , John Jairo Molina , Ryoichi Yamamoto , Takashi Taniguchi

Although human beings have known about the phenomenon of "flocking"- that is, the coherent movement of large numbers of creatures (flocks of birds, schools of fish, herds of woolly mammoths, etc.)- since prehistoric times, it is only in the…

Soft Condensed Matter · Physics 2018-12-04 John Toner

Collective behaviour in living systems is observed across many scales, from bacteria to insects, to fish shoals. Zebrafish have emerged as a model system amenable to laboratory study. Here we report a three-dimensional study of the…

Statistical Mechanics · Physics 2024-05-16 Yushi Yang , Francesco Turci , Erika Kague , Chrissy L. Hammond , John Russo , C. Patrick Royall

We construct the hydrodynamic theory of coherent collective motion ("flocking") at a solid-liquid interface. The polar order parameter and concentration of a collection of "active" (self-propelled) particles at a planar interface between a…

Soft Condensed Matter · Physics 2022-01-05 Niladri Sarkar , Abhik Basu , John Toner

We numerically examine a bidisperse system of active and passive particles coupled to a resource substrate. The active particles deplete the resource at a fixed rate and move toward regions with higher resources, while all of the particles…

Soft Condensed Matter · Physics 2024-04-23 L. Varga , A. Libal , C. Reichhardt , C. J. O. Reichhardt

Natural flocks (aligned) and swarms (non-aligned) both exhibit features of near-criticality, challenging their treatment as two ends of the same phase transition. We present a model for the aggregation of active individuals, in which their…

Adaptation and Self-Organizing Systems · Physics 2025-11-26 Joao Lizárraga , Marcus de Aguiar

Animals typically forage in groups. Social foraging can help animals avoid predation and decrease their uncertainty about the richness of food resources. Despite this, theoretical mechanistic models of patch foraging have overwhelmingly…

Populations and Evolution · Quantitative Biology 2022-02-14 Subekshya Bidari , Ahmed El Hady , Jacob Davidson , Zachary P Kilpatrick

The spontaneous emergence of collective motion patterns is usually associated with the presence of a velocity alignment mechanism that mediates the interactions among the moving individuals. Despite of this widespread view, it has been…

Biological Physics · Physics 2019-12-18 Fernando Peruani