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The cytoskeletal component actomyosin is a canonical example of active matter since the powerstroke cycle locally converts chemical energy in the form of adenoside triphosphate (ATP) into mechanical work for remodelling. Observing myosin II…

Biological Physics · Physics 2024-12-05 Sami C. Al-Izzi , Sedigheh Ghanbarzadeh Nodehi , Darius V. Köster , Richard G. Morris

Oscillatory behaviour in force-generating systems is a pervasive phenomenon in cell biology. In this work, we investigate how oscillations in the actomyosin cytoskeleton drive cell shape changes during the process of Dorsal Closure, a…

Cell Behavior · Quantitative Biology 2015-06-18 Pedro F. Machado , Guy B. Blanchard , Julia Duque , Nicole Gorfinkiel

Contractility in animal cells is often generated by molecular motors such as myosin, which require polar substrates for their function. Motivated by recent experimental evidence of motor-independent contractility, we propose a robust…

Soft Condensed Matter · Physics 2022-05-04 Sihan Chen , Tomer Markovich , Fred C. MacKintosh

Forces in the actin cytoskeleton are generated by small groups of non-processive myosin II motors for which stochastic effects are highly relevant. Using a crossbridge model with the assumptions of fast powerstroke kinetics and equal load…

Subcellular Processes · Quantitative Biology 2015-06-04 Thorsten Erdmann , Ulrich S. Schwarz

Cellular cargo can be bound to cytoskeletal filaments by one or multiple active or passive molecular motors. Recent experiments have shown that the presence of auxiliary, nondriving motors, results in an enhanced processivity of the cargo,…

Biomolecules · Quantitative Biology 2019-03-27 Filippo Posta , Maria R. D'Orsogna , Tom Chou

Cytoskeletal networks are the main actuators of cellular mechanics, and a foundational example for active matter physics. In cytoskeletal networks, motion is generated on small scales by filaments that push and pull on each other via…

Soft Condensed Matter · Physics 2022-10-05 Sebastian Fürthauer , Michael J. Shelley

Current understanding of how contractility emerges in disordered actomyosin networks of non-muscle cells is still largely based on the intuition derived from earlier works on muscle contractility. This view, however, largely overlooks the…

Biological Physics · Physics 2018-04-11 James E. Komianos , Garegin A. Papoian

The synthetic actin network demands great interest as bio-material due to its soft and wet nature that mimic many biological scaffolding structures. The study of these structures can contribute to a better understanding of this new…

Soft Condensed Matter · Physics 2017-02-10 Horacio López-Menéndez , José Félix Rodríguez

Motor-driven cytoskeletal remodeling in cellular systems can often be accompanied by a diffusive-like effect at local scales, but distinguishing the contributions of the ordering process, such as active contraction of a network, from this…

Biological Physics · Physics 2024-08-22 Soichi Hirokawa , Heun Jin Lee , Rachel A Banks , Ana I Duarte , Bibi Najma , Matt Thomson , Rob Phillips

The actin cytoskeleton of cells is in continuous motion due to both polymerization of new filaments and their contraction by myosin II molecular motors. Through adhesion to the substrate, such intracellular flow can be converted into cell…

Subcellular Processes · Quantitative Biology 2021-08-11 Oliver M. Drozdowski , Falko Ziebert , Ulrich S. Schwarz

The actomyosin cortex is an active material that provides animal cells with a strong but flexible exterior, whose mechanics, including non-Gaussian fluctuations and occasional large displacements or cytoquakes, have defied explanation. We…

It is well known that many biochemical processes in the cell such as gene regulation, growth signals and activation of ion channels, rely on mechanical stimuli. However, the mechanism by which mechanical signals propagate through cells is…

Soft Condensed Matter · Physics 2017-04-05 Adar Sonn-Segev , Anne Bernheim-Groswasser , Yael Roichman

The growth of actin filament networks is a fundamental biological process that drives a variety of cellular and intracellular motions. During motility, eukaryotic cells and intracellular pathogens are propelled by actin networks organized…

Biological Physics · Physics 2009-11-13 Joshua W. Shaevitz , Daniel A. Fletcher

The emergence of macroscopic order and patterns is a central paradigm in systems of (self-)propelled agents, and a key component in the structuring of many biological systems.The relationships between the ordering process and the underlying…

Soft Condensed Matter · Physics 2020-12-02 Jonas Denk , Erwin Frey

Mechanically induced unfolding of passive crosslinkers is a fundamental biological phenomenon encountered across the scales from individual macro-molecules to cytoskeletal actin networks. In this paper we study a conceptual model of…

Biological Physics · Physics 2015-01-08 M Caruel , J. -M Allain , L Truskinovsky

Biological activity gives rise to non-equilibrium fluctuations in the cytoplasm of cells; however, there are few methods to directly measure these fluctuations. Using a reconstituted actin cytoskeleton, we show that the bending dynamics of…

Soft Condensed Matter · Physics 2009-11-13 C. P. Brangwynne , G. H. Koenderink , F. C. MacKintosh , D. A. Weitz

We propose a novel mechanism of cell motility, which relies on the coupling of actin polymerization at the cell membrane to geometric confinement. We consider a polymerizing viscoelastic cytoskeletal gel confined in a narrow channel, and…

Soft Condensed Matter · Physics 2009-02-13 R. J. Hawkins , M. Piel , G. Faure-Andre , A. M. Lennon-Dumenil , J. F. Joanny , J. Prost , R. Voituriez

Many biological processes are supported by special molecules, called motor proteins or molecular motors, that transport cellular cargoes along linear protein filaments and can reversibly associate to their tracks. Stimulated by these…

Statistical Mechanics · Physics 2021-11-17 Akriti Jindal , Anatoly B. Kolomeisky , Arvind Kumar Gupta

Biochemistry and mechanics are closely coupled in cell adhesion. At sites of cell-matrix adhesion, mechanical force triggers signaling through the Rho-pathway, which leads to structural reinforcement and increased contractility in the actin…

Subcellular Processes · Quantitative Biology 2010-02-24 A. Besser , U. S. Schwarz

The initiation of directional cell motion requires symmetry breaking that can happen both with or without external stimuli. During cell crawling, forces generated by the cytoskeleton and their transmission through mechanosensitive adhesions…

Cell Behavior · Quantitative Biology 2023-06-02 Yuzhu Chen , David Saintillan , Padmini Rangamani