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Biological evolution can be conceptualized as a search process in the space of gene sequences guided by the fitness landscape, a mapping that assigns a measure of reproductive value to each genotype. Here we discuss probabilistic models of…

Populations and Evolution · Quantitative Biology 2024-04-10 Joachim Krug , Daniel Oros

The fitness landscape encodes the mapping of genotypes to fitness and provides a succinct representation of possible trajectories followed by an evolving population. Evolutionary accessibility is quantified by the existence of…

Populations and Evolution · Quantitative Biology 2021-06-30 Joachim Krug

The adaptive evolution of a population under the influence of mutation and selection is strongly influenced by the structure of the underlying fitness landscape, which encodes the interactions between mutations at different genetic loci.…

Populations and Evolution · Quantitative Biology 2015-06-04 Jasper Franke , Joachim Krug

A fitness landscape is a mapping from the space of genetic sequences, which is modeled here as a binary hypercube of dimension $L$, to the real numbers. We consider random models of fitness landscapes, where fitness values are assigned…

Populations and Evolution · Quantitative Biology 2015-06-16 Benjamin Schmiegelt , Joachim Krug

These notes introduce probabilistic landscape models defined on high-dimensional discrete sequence spaces. The models are motivated primarily by fitness landscapes in evolutionary biology, but links to statistical physics and computer…

Populations and Evolution · Quantitative Biology 2025-12-24 Sakshi Pahujani , Joachim Krug

All possible phenotypes are not equally accessible to evolving populations. In fact, only phenotypes of large size, i.e. those resulting from many different genotypes, are found in populations of sequences, presumably because they are…

Populations and Evolution · Quantitative Biology 2023-08-16 Pablo Catalán , Juan Antonio García-Martín , Jacobo Aguirre , José A. Cuesta , Susanna Manrubia

The fitness landscape - the mapping between genotypes and fitness - determines properties of the process of adaptation. Several small genetic fitness landscapes have recently been built by selecting a handful of beneficial mutations and…

Populations and Evolution · Quantitative Biology 2014-05-15 François Blanquart , Guillaume Achaz , Thomas Bataillon , Olivier Tenaillon

A fitness landscape is a mapping from a space of discrete genotypes to the real numbers. A path in a fitness landscape is a sequence of genotypes connected by single mutational steps. Such a path is said to be accessible if the fitness…

Populations and Evolution · Quantitative Biology 2023-02-21 Benjamin Schmiegelt , Joachim Krug

Molecular evolution is often conceptualised as adaptive walks on rugged fitness landscapes, driven by mutations and constrained by incremental fitness selection. It is well known that epistasis shapes the ruggedness of the landscape's…

Populations and Evolution · Quantitative Biology 2022-11-03 Leonardo Trujillo , Paul Banse , Guillaume Beslon

The mechanisms by which adaptive phenotypes spread within an evolving population after their emergence are understood fairly well. Much less is known about the factors that influence the evolutionary accessibility of such phenotypes, a…

Populations and Evolution · Quantitative Biology 2015-05-13 Wilfred Ndifon , Joshua B. Plotkin , Jonathan Dushoff

A common view in evolutionary biology is that mutation rates are minimised. However, studies in combinatorial optimisation and search have shown a clear advantage of using variable mutation rates as a control parameter to optimise the…

Populations and Evolution · Quantitative Biology 2019-08-27 Roman V. Belavkin , Alastair Channon , Elizabeth Aston , John Aston , Rok Krasovec , Christopher G. Knight

The tempo and mode of an adaptive process is strongly determined by the structure of the fitness landscape that underlies it. In order to be able to predict evolutionary outcomes (even on the short term), we must know more about the nature…

Populations and Evolution · Quantitative Biology 2013-02-28 Bjørn Østman , Christoph Adami

Epistatic interactions between mutations add substantial complexity to adaptive landscapes, and are often thought of as detrimental to our ability to predict evolution. Yet, patterns of global epistasis, in which the fitness effect of a…

Populations and Evolution · Quantitative Biology 2022-10-10 Juan Diaz-Colunga , Abigail Skwara , Karna Gowda , Ramon Diaz-Uriarte , Mikhail Tikhonov , Djordje Bajic , Alvaro Sanchez

Evolutionary adaptation is the process that increases the fit of a population to the fitness landscape it inhabits. As a consequence, evolutionary dynamics is shaped, constrained, and channeled, by that fitness landscape. Much work has been…

Populations and Evolution · Quantitative Biology 2010-12-17 Bjørn Østman , Arend Hintze , Christoph Adami

This is an introductory review of deterministic mutation-selection models for asexual populations (i.e., quasispecies theory) and related topics. First, the basic concepts of fitness, mutations, and sequence space are introduced. Different…

Populations and Evolution · Quantitative Biology 2007-07-26 Kavita Jain , Joachim Krug

We study biological evolution in a high-dimensional genotype space in the regime of rare mutations and strong selection. The population performs an uphill walk which terminates at local fitness maxima. Assigning fitness randomly to…

Populations and Evolution · Quantitative Biology 2015-05-28 Johannes Neidhart , Joachim Krug

Mutational robustness quantifies the effect of random mutations on fitness. When mutational robustness is high, most mutations do not change fitness or have only a minor effect on it. From the point of view of fitness landscapes, robust…

Populations and Evolution · Quantitative Biology 2019-10-22 Alexander Klug , Su-Chan Park , Joachim Krug

We introduce a new model of evolution on a fitness landscape possessing a tunable degree of neutrality. The model allows us to study the general properties of molecular species undergoing neutral evolution. We find that a number of…

adap-org · Physics 2007-05-23 M. E. J. Newman , Robin Engelhardt

Fitness landscapes are mappings between genotypes, phenotypes, and fitness that shape evolution. In recent years, empirical work and theoretical models have greatly advanced our understanding of how populations navigate rugged fitness…

Populations and Evolution · Quantitative Biology 2026-04-21 Malvika Srivastava , Claudia Bank , Joachim Krug , Suman G. Das

The fitness landscape metaphor plays a central role on the modeling of optimizing principles in many research fields, ranging from evolutionary biology, where it was first introduced, to management research. Here we consider the ensemble of…

Populations and Evolution · Quantitative Biology 2019-01-30 Paulo R. A. Campos , José F. Fontanari
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