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We study a reconstituted composite system consisting of an active microtubule network interdigitated with a passive network of entangled F-actin filaments. Increasing viscoelasticity of the F-actin network controls the emergent dynamics,…

Soft Condensed Matter · Physics 2022-02-16 John Berezney , Bruce L. Goode , Seth Fraden , Zvonimir Dogic

Actin is a key component of the cytoskeleton, which plays central roles in cell motility, division, growth, and tensile strength. To enable this wide range of transient mechanical processes and properties, networks of actin filaments…

Centering and decentering of cellular components is essential for internal organization of cells and their ability to perform basic cellular functions such as division and motility. How cells achieve proper localization of their components…

Eukaryotic cells and intracellular pathogens such as bacteria or viruses utilize the actin polymerization machinery to propel themselves forward. Thereby, the onset of motion and choice of direction may be the result of a spontaneous…

Soft Condensed Matter · Physics 2009-09-04 Karin John , Denis Caillerie , Philippe Peyla , Mourad Ismail , Annie Raoult , Jacques Prost

Inspired by recent experiments of cells accumulating on anisotropic substrates, we study a two-dimensional, compressible, isotropic, active fluid in the presence of anisotropic friction. We find that regions of anisotropic friction that are…

Soft Condensed Matter · Physics 2026-03-04 Cody D. Schimming , Brian A. Camley

Transition from random to persistent cell motility requires spatiotemporal organization of the cytoskeleton and focal adhesions. The influence of these two structures on cell steering can also be gleaned from trypsin de-adhesion…

Cell Behavior · Quantitative Biology 2013-10-29 Soumya S S , Subodh Kolwankar , Edna George , Santanu K. Basu , Shamik Sen , Mandar M. Inamdar

The dynamics of a membrane coupled to an active fluid on top of a substrate is considered theoretically. It is assumed that the director field of the active fluid has rotational symmetry in the membrane plane. This situation is likely to be…

Soft Condensed Matter · Physics 2020-04-08 Chia-Chun Liang , Kento Yasuda , Shigeyuki Komura , Kuo-An Wu , Hsuan-Yi Chen

The tubulin-like protein FtsZ is crucial for cytokinesis in bacteria and many archaea, forming a ring-shaped structure called the Z-ring at the site of cell division. Despite extensive research, the self-assembly of Z-rings is not entirely…

Biological Physics · Physics 2025-02-17 Rajneesh Kumar , Ramanujam Srinivasan , Debasish Chaudhuri

Actin networks, acting as an engine pushing against an external load, are fundamentally important to cell motility. A measure of the effectiveness of an engine is the velocity the engine is able to produce at a given force, the…

Subcellular Processes · Quantitative Biology 2011-11-30 Daniel B. Smith , Jian Liu

We investigate the force generation by polymerizing bundles of filaments, which form because of short-range attractive filament interactions. We show that bundles can generate forces by a zipping mechanism, which is not limited by buckling…

Soft Condensed Matter · Physics 2012-05-02 Torsten Kuehne , Reinhard Lipowsky , Jan Kierfeld

The actin cytoskeleton in living cells has many types of crosslinkers. The mechanical interplay between these different crosslinker types is an open issue in cytoskeletal mechanics. We develop a framework to study the cooperativity and…

Soft Condensed Matter · Physics 2015-05-28 Moumita Das , D. A. Quint , J. M. Schwarz

Cytoskeletal filaments transported by surface immobilized molecular motors with one end pinned to the surface have been observed to spiral in a myosin-driven actin 'gliding assay'. The radius of the spiral was shown to scale with motor…

Soft Condensed Matter · Physics 2025-08-15 Aman Soni , Shivani A. Yadav , Chaitanya A. Athale

We apply a recently developed model of cytoskeletal force generation to study a cell intrinsic contractility, as well as its response to external loading. The model is based on a non-equilibrium thermodynamic treatment of the…

Subcellular Processes · Quantitative Biology 2015-06-19 Mirko Maraldi , Clara Valero , Krishna Garikipati

Cross-linked polymer networks with orientational order constitute a wide class of soft materials and are relevant to biological systems (e.g., F-actin bundles). We analytically study the nonlinear force-extension relation of an array of…

Disordered Systems and Neural Networks · Physics 2015-06-05 Panayotis Benetatos , Stephan Ulrich , Annette Zippelius

A generalized theory of the self-limiting assembly of twisted bundles of filaments and columns is presented. Bundles and fibers form in a broad variety of supramolecular systems, from biological to synthetic materials. A widely-invoked…

Soft Condensed Matter · Physics 2023-08-08 Gregory M. Grason

Robust self-organization of subcellular structures is a key principle governing the dynamics and evolution of cellular life. In fission yeast cells undergoing division, the mitotic spindle spontaneously emerges from the interaction of…

Soft Condensed Matter · Physics 2024-06-03 David A. Head , W. J. Briels , Gerhard Gompper

Left-right axis specification establishes embryonic laterality through asymmetric signaling cascades originating at the cellular scale. We previously reported the presence of a directionality bias in confined pairs of endothelial (and…

Cell Behavior · Quantitative Biology 2026-03-31 Egun Im , Ghina Badih , Laetitia Kurzawa , Andreas Buttenschön , Calina Copos

Recent advances in microscopy techniques has uncovered unique aspects of flagella-driven motility in bacteria. A remarkable example is the discovery of flagellar wrapping, a phenomenon whereby a bacterium wraps its flagellum (or flagellar…

Soft Condensed Matter · Physics 2025-04-22 Takuro Kataoka , Taiju Yoneda , Daisuke Nakane , Hirofumi Wada

Non-processive molecular motors have to work together in ensembles in order to generate appreciable levels of force or movement. In skeletal muscle, for example, hundreds of myosin II molecules cooperate in thick filaments. In non-muscle…

Biological Physics · Physics 2015-06-16 Thorsten Erdmann , Philipp J. Albert , Ulrich S. Schwarz

We investigate the formation of helical multifilament bundles and the torque required to achieve them as a function of applied twist. Hyperelastic filaments with circular cross sections are mounted parallel in a uniform circle onto…

Soft Condensed Matter · Physics 2026-04-01 Amit Dawadi , Animesh Biswas , Julien Chopin , Arshad Kudrolli