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The behaviour of microscopic swimmers has previously been explored near large scale confining geometries and in the presence of very small-scale surface roughness. Here we consider an intermediate case of how a simple microswimmer, the…

Fluid Dynamics · Physics 2021-09-14 Kenta Ishimoto , Eamonn A. Gaffney , David J. Smith

Hydrodynamic interactions are crucial for determining the cooperative behavior of microswimmers at low Reynolds numbers. Here we provide a comprehensive analysis of the scaling and strength of the interactions in the case of a pair of…

Soft Condensed Matter · Physics 2021-02-11 Sebastian Ziegler , Thomas Scheel , Maxime Hubert , Jens Harting , Ana-Sunčana Smith

We theoretically and computationally study the low-Reynolds-number hydrodynamics of a linear active microswimmer surfing on a compressible thin fluid layer characterized by an odd viscosity. Since the underlying three-dimensional fluid is…

Soft Condensed Matter · Physics 2023-10-25 Yuto Hosaka , Ramin Golestanian , Abdallah Daddi-Moussa-Ider

The majority of studies on self-propelled particles and microswimmers concentrates on objects that do not feature a deterministic bending of their trajectory. However, perfect axial symmetry is hardly found in reality, and shape-asymmetric…

Statistical Mechanics · Physics 2018-07-18 Christian Hoell , Hartmut Löwen , Andreas M. Menzel

Motility is an essential factor for an organism's survival and diversification. With the advent of novel single-cell technologies, analytical frameworks and theoretical methods, we can begin to probe the complex lives of microscopic motile…

Both legged locomotion and slithering motions typically utilize periodic gaits -- repeating cycles of body shape change that produce a net motion through the world. Legged locomotion can be viewed from the perspective of piece-wise contact…

Robotics · Computer Science 2022-05-03 Shai Revzen , Dan Zhao , Glenna Clifton , Brian Bittner , Nick Gravish

We show using theory and experiments that a small particle moving along an elastic membrane through a viscous fluid is repelled from the membrane due to hydro-elastic forces. The viscous stress field produces an elastic disturbance leading…

Soft Condensed Matter · Physics 2018-02-23 Bhargav Rallabandi , Naomi Oppenheimer , Matan Yah Ben Zion , Howard A. Stone

We study a random walk driven by a particle system from a generic class, and establish a law of large numbers for the walk for almost all densities of the environment. To do so, we exploit the finite-ranged approximations of the environment…

Probability · Mathematics 2026-05-27 Guillaume Conchon--Kerjan , Toril Palaniappan

We experimentally study a scallop-like swimmer with reciprocally flapping wings in a nearly frictionless, cohesive granular medium consisting of hydrogel spheres. Significant locomotion is found when the swimmer's flapping frequency matches…

Soft Condensed Matter · Physics 2026-04-29 Hongyi Xiao , Jing Wang , Achim Sack , Ralf Stannarius , Thorsten Pöschel

The geometric phase techniques for swimming in viscous flows express the net displacement of a swimmer as a path integral of a field in configuration space. This representation can be transformed into an area integral for simple swimmers…

Fluid Dynamics · Physics 2021-04-13 Lyndon Koens , Eric Lauga

Motivated by recent advances in vesicle engineering, we consider theoretically the locomotion of shape-changing bilayer vesicles at low Reynolds number. By modulating their volume and membrane composition, the vesicles can be made to change…

Biological Physics · Physics 2010-04-09 Arthur A. Evans , Saverio E. Spagnolie , Eric Lauga

Simple, linear equations relate microscopic swimmers to the corresponding gliders and pumps. They have the following set of consequences: The swimming velocity of free swimmers can be inferred from the force on the tethered swimmer and vice…

Fluid Dynamics · Physics 2010-07-28 J. E. Avron , O. Raz

We provide exact solutions of the Stokes equations for a squirming sphere close to a no-slip surface, both planar and spherical, and for the interactions between two squirmers, in three dimensions. These allow the hydrodynamic interactions…

Fluid Dynamics · Physics 2017-04-05 Dario Papavassiliou , Gareth P. Alexander

The effect of rotational constraint on the properties of lattice models like the self-avoiding walk, lattice animals and percolation is discussed. The results obtained so far, using a variety of exact and approximate techniques, are…

Statistical Mechanics · Physics 2008-02-03 Indrani Bose

The persistent dynamics of active particles makes them explore extended portions of an obstacle's boundary during collisions. From impact to escape, the net applied forces depend on the curvature of the wall and increase in the presence of…

Unlike macroscopic swimmers, microswimmers operate in a low-Reynolds-number regime dominated by viscous forces. This paper investigates the controllability of a magnetic microswimmer composed of a spherical magnetic head and an elastic,…

Optimization and Control · Mathematics 2025-11-05 Lucas Palazzolo , Mickaël Binois , Laëtitia Giraldi

We use particle simulations to reveal two distinct propulsion mechanisms for a scallop-like swimmer to locomote itself in granular media by reciprocally flapping its wings. Based on the discrete element method, we examine the kinematics and…

Soft Condensed Matter · Physics 2025-10-28 Amir Nazemi , Hongyi Xiao

Many micro-swimmers propel themselves by rotating micro-cylindrical organelles such as flagella or cilia. These cylindrical organelles almost never live in free space, yet their motions in a confining geometry can be counter-intuitive. For…

Fluid Dynamics · Physics 2023-12-25 Hanliang Guo , Yi Man , Hai Zhu

The macroscopic behavior of dense suspensions of neutrally-buoyant spheres in turbulent plane channel flow is examined. We show that particles larger than the smallest turbulence scales cause the suspension to deviate from the continuum…

Fluid Dynamics · Physics 2016-09-28 Pedro Costa , Francesco Picano , Luca Brandt , Wim-Paul Breugem

This paper focuses on the mechanism of propulsion of a Purcell swimmer whose segments are magnetized and react to an external magnetic field applied into the fluid. By an asymptotic analysis, we prove that it is possible to steer the…

Dynamical Systems · Mathematics 2016-11-08 François Alouges , Antonio Desimone , Laetitia Giraldi , Marta Zoppello