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The underlying mechanisms and physics of catalytic Janus microswimmers is highly complex, requiring details of the associated phoretic fields and the physiochemical properties of catalyst, particle, boundaries, and the fuel used. Therefore,…

We use particle simulations to reveal two distinct propulsion mechanisms for a scallop-like swimmer to locomote itself in granular media by reciprocally flapping its wings. Based on the discrete element method, we examine the kinematics and…

Soft Condensed Matter · Physics 2025-10-28 Amir Nazemi , Hongyi Xiao

It has been proposed that biologically generated turbulence plays an important role in material transport and ocean mixing. Both experimental and numerical studies have reported evidence of the non-negligible mixing by moderate Reynolds…

Fluid Dynamics · Physics 2023-10-06 Xinyu Si , Lei Fang

Cooperative motion in biological microswimmers is crucial for their survival as it facilitates adhesion to surfaces, formation of hierarchical colonies, efficient motion, and enhanced access to nutrients. Synthetic microswimmers currently…

Soft Condensed Matter · Physics 2022-04-12 Stefania Ketzetzi , Melissa Rinaldin , Pim Dröge , Joost de Graaf , Daniela J. Kraft

The flow-driven transport of interacting micron-sized particles occurs in many soft matter systems spanning from the translocation of proteins to moving emulsions in microfluidic devices. Here we combine experiments and theory to…

Soft Condensed Matter · Physics 2022-11-16 Dominik Lips , Eric Cereceda-López , Antonio Ortiz-Ambriz , Pietro Tierno , Artem Ryabov , Philipp Maass

In their search for metabolic resources microbes swim through viscous environments that present physical anisotropies, including steric obstacles across a wide range of sizes. Hydrodynamic forces are known to significantly alter swimmer…

Soft Condensed Matter · Physics 2020-07-03 Kentaro Hoeger , Tristan Ursell

We analyze the dynamics of a microswimmer in pressure-driven Poiseuille flow, where fluid inertia is small but non-negligible. Using perturbation theory and the reciprocal theorem, we show that in addition to the classical inertial lift of…

Soft Condensed Matter · Physics 2021-09-21 Akash Choudhary , Subhechchha Paul , Felix Rühle , Holger Stark

Despite their vast morphological diversity, many invertebrates have similar larval forms characterized by ciliary bands, innervated arrays of beating cilia that facilitate swimming and feeding. Hydrodynamics suggests that these bands should…

Fluid Dynamics · Physics 2018-04-25 William Gilpin , Vivek N. Prakash , Manu Prakash

The hydrodynamic interactions of a suspension of self-propelled particles are studied using a direct numerical simulation method which simultaneously solves for the host fluid and the swimming particles. A modified version of the "Smoothed…

Soft Condensed Matter · Physics 2013-01-15 John J. Molina , Yasuya Nakayama , Ryoichi Yamamoto

Hydrodynamic interactions between fluid-dispersed particles are ubiquitous in soft matter and biological systems and they give rise to intriguing collective phenomena. While it was reported that these interactions can facilitate…

Soft Condensed Matter · Physics 2022-08-29 Eric Cereceda-López , Dominik Lips , Antonio Ortiz-Ambriz , Artem Ryabov , Philipp Maass , Pietro Tierno

A theory is presented for wave-driven propulsion of floating bodies driven into oscillation at the fluid interface. By coupling the equations of motion of the body to a quasi-potential flow model of the fluid, we derive expressions for the…

Fluid Dynamics · Physics 2023-10-20 GP Benham , O Devauchelle , SJ Thomson

For natural microswimmers, the interplay of swimming activity and external flow can promote robust motion, e.g. propulsion against ("upstream rheotaxis") or perpendicular to the direction of flow. These effects are generally attributed to…

Soft Condensed Matter · Physics 2018-01-30 Jaideep Katuri , William E. Uspal , Juliane Simmchen , Albert Miguel-López , Samuel Sánchez

In this paper we study crawling locomotion based on directional frictional interactions, namely, frictional forces that are sensitive to the sign of the sliding velocity. Surface interactions of this type are common in biology, where they…

Mathematical Physics · Physics 2015-03-31 Paolo Gidoni , Giovanni Noselli , Antonio DeSimone

The rectilinear "drift" of particles in a hydrodynamic drift ratchet arises from a combination of diffusive motion and particle-wall hydrodynamic interactions, and is therefore dependent on particle diffusivity, particle size, the amplitude…

Marine plankton play a crucial role in carbon storage, oxygen production, global climate, and ecosystem function. Planktonic ecosystems are embedded in a Lagrangian patches of water that are continuously moving, stretching, and diluting.…

Populations and Evolution · Quantitative Biology 2023-01-30 Enrico Ser-Giacomi , Ricardo Martinez-Garcia , Stephanie Dutkiewicz , Michael J. Follows

We study the problem of sinking particles in a realistic oceanic flow, with major energetic structures in the mesoscale, focussing in the range of particle sizes and densities appropriate for marine biogenic particles. Our aim is to unify…

Atmospheric and Oceanic Physics · Physics 2017-07-03 Pedro Monroy , Emilio Hernández-García , Vincent Rossi , Cristóbal López

Microswimmers, especially in theoretical treatments, are generally taken to be completely inertia-free, since inertial effects on their motion are typically small and assuming their absence simplifies the problem considerably. Yet in nature…

Soft Condensed Matter · Physics 2016-11-08 Jayant Pande , Kristina Pickl , Oleg Trosman , Ulrich Rüde , Ana-Sunčana Smith

We analyze a minimal model for a rigid spherical microswimmer and explore the consequences of its extended surface on the interplay between its self-propulsion and flow properties. The model is the first order representation of…

Soft Condensed Matter · Physics 2017-12-06 Tapan Chandra Adhyapak , Sara Jabbari-Farouji

We numerically investigate the motion of active artificial microswimmers diffusing in a fuel concentration gradient. We observe that, in the steady state, their probability density accumulates in the low-concentration regions, whereas a…

Soft Condensed Matter · Physics 2025-08-18 Pulak K. Ghosh , Yunyun Li , Fabio Marchesoni , Franco Nori

Mesozooplankton aggregations undergoing vertical migrations in the upper ocean have been hypothesized to have an important role in the redistribution of carbon, nutrients, and oxygen via biogenic hydrodynamic transport (BHT). While…

Fluid Dynamics · Physics 2026-03-04 Yunxing Su , Rui Zhu , Eckart Meiburg , Monica M. Wilhelmus