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The inner ear achieves a wide dynamic range of responsiveness by mechanically amplifying weak sounds. The enormous mechanical gain reported for the mammalian cochlea, which exceeds a factor of 4,000, poses a challenge for theory. Here we…

Tissues and Organs · Quantitative Biology 2010-09-13 Tobias Reichenbach , A. J. Hudspeth

The extraordinary sensitivity of the mammalian inner ear has captivated scientists for decades, largely due to the crucial role played by the outer hair cells (OHCs) and their unique electromotile properties. Typically arranged in three…

Biological Physics · Physics 2023-11-16 Alessandro Altoè , Christopher A Shera

Most sounds of interest consist of complex, time-dependent admixtures of tones of diverse frequencies and variable amplitudes. To detect and process these signals, the ear employs a highly nonlinear, adaptive, real-time spectral analyzer:…

Neurons and Cognition · Quantitative Biology 2014-08-12 T. Reichenbach , A. J. Hudspeth

The ability of the mammalian ear in processing high frequency sounds, up to $\sim$100 kHz, is based on the capability of outer hair cells (OHCs) responding to stimulation at high frequencies. These cells show a unique motility in their cell…

Biological Physics · Physics 2017-09-22 Kuni H Iwasa

A mathematical model describing the coupling between two independent amplification mechanisms in auditory hair cells is proposed and analyzed. Hair cells are cells in the inner ear responsible for translating sound-induced mechanical…

Pattern Formation and Solitons · Physics 2009-11-13 K. A. Montgomery , M. Silber , S. A. Solla

Hair cells, the sensory receptors of the internal ear, subserve different functions in various receptor organs: they detect oscillatory stimuli in the auditory system, but transduce constant and step stimuli in the vestibular and…

Neurons and Cognition · Quantitative Biology 2017-03-29 Joshua D. Salvi , Daibhid O Maoileidigh , Brian A. Fabella , Melanie Tobin , A. J. Hudspeth

The cochlea or inner ear has a remarkable ability to amplify sound signals. This is understood to derive at least in part from some active process that magnifies vibrations of the basilar membrane (BM) and the cochlear partition in which it…

Biological Physics · Physics 2015-07-08 William Ko , John M. Stockie

High auditory sensitivity, sharp frequency selectivity, and otoacoustic emissions are signatures of active amplification of the cochlea. The human ear can also detect very large amplitude sound without being damaged as long as the exposed…

Biological Physics · Physics 2008-10-01 Yong Zhang , Chul Koo Kim , Kong-Ju-Bock Lee , Youngah Park

The hair cells of the vertebrate inner ear convert mechanical stimuli to electrical signals. Two adaptation mechanisms are known to modify the ionic current flowing through the transduction channels of the hair bundles: a rapid process…

Biological Physics · Physics 2009-11-10 Andrej Vilfan , Thomas Duke

We introduce the concept of self-tuned criticality as a general mechanism for signal detection in sensory systems. In the case of hearing, we argue that active amplification of faint sounds is provided by a dynamical system which is…

Biological Physics · Physics 2009-11-06 Sebastien Camalet , Thomas Duke , Frank Julicher , Jacques Prost

Hair cells of the auditory and vestibular systems display astonishing sensitivity, frequency selectivity, and temporal resolution to external signals. These specialized cells utilize an internal active amplifier to achieve highly sensitive…

Neurons and Cognition · Quantitative Biology 2021-03-31 Justin Faber , Hancheng Li , Dolores Bozovic

Electromotility of outer hair cells (OHCs) has been extensively studied with in vitro experiments because of its physiological significance in the cochlear amplifier, which provides the exquisite sensitivity and frequency selectivity of the…

Biological Physics · Physics 2016-12-08 Kuni H Iwasa

The outer hair cell (OHC) of the organ of Corti underlies a mechanically based process that enhances hearing, termed cochlear amplification. The cell possesses a unique motor protein, prestin, which senses voltage and consequently changes…

Biological Physics · Physics 2020-01-29 Joseph Santos-Sacchi , Kuni Iwasa , Winston Tan

Outer hair cells (OHCs) are essential for the sensitivity and frequency specificity of the mammalian ear. To perform this function, OHCs need to amplify the motion of the basilar membrane (BM), which is much stiffer than themselves. If OHCs…

Biological Physics · Physics 2026-02-05 Kuni H. Iwasa

Sound produces surface waves along the cochlea's basilar membrane. To achieve the ear's astonishing frequency resolution and sensitivity to faint sounds, dissipation in the cochlea must be canceled via active processes in hair cells,…

Biological Physics · Physics 2025-04-23 Asheesh S. Momi , Michael C. Abbott , Julian Rubinfien , Benjamin B. Machta , Isabella R. Graf

Hair cells of the auditory and vestibular systems are capable of detecting sounds that induce sub-nanometer vibrations of the hair bundle, below the stochastic noise levels of the surrounding fluid. Hair bundles of certain species are also…

Neurons and Cognition · Quantitative Biology 2019-05-23 Justin Faber , Dolores Bozovic

An active dissipative process organizes auditory frequency analysis in the mammalian cochlea. A minimal active beam model reveals that a spatially varying viscous coupling operator, $\partial_{xx}\kappa\partial_{xx}$, generates dissipative…

Biological Physics · Physics 2026-02-06 Yasuki Murakami

Sensory hair cells in auditory and vestibular organs rely on active mechanisms to achieve high sensitivity and frequency selectivity. Recent experimental studies have documented self-sustained oscillations in hair cells of lower vertebrates…

Biological Physics · Physics 2012-09-28 Rami Amro , Alexander B. Neiman

Cochlear outer hair cells (OHCs) have two mechanosensitive elements: the hair bundle with mechanotrasducer channels and the piezoelectric lateral wall of the cell body. The present report examines how these elements interact with each other…

Biological Physics · Physics 2025-09-09 Kuni H Iwasa

Hair cells actively drive oscillations of their mechanosensitive organelles--the hair bundles that enable hearing and balance sensing in vertebrates. Why and how some hair cells expend energy by sustaining this oscillatory motion in order…

Statistical Mechanics · Physics 2026-04-02 Yanathip Thipmaungprom , Laila Saliekh , Rodrigo Alonso , Édgar Roldán , Florian Berger , Roman Belousov
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