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We show that a two-dimensional system of flocking microswimmers interacting hydrodynamically can be expressed using a Hamiltonian formalism. The Hamiltonian depends strictly on the angles between the particles and their swimming…

Soft Condensed Matter · Physics 2023-05-23 Yuval Shoham , Naomi Oppenheimer

Several micro-organisms, such as bacteria, algae, or spermatozoa, use flagella or cilia to swim in a fluid, while many other micro-organisms instead use ample shape deformation, described as amoeboid, to propel themselves by either crawling…

Biological Physics · Physics 2016-09-19 Hao Wu , A. Farutin , W. -F. Hu , M. Thiébaud , S. Rafaï , P. Peyla , M. -C. Lai , C. Misbah

We develop a numerical framework to simulate the locomotion of a flagellated bacterium with a spheroidal head (such as Escherichia coli) in biological fluids like mucus, which are entangled polymer solutions exhibiting elasto-viscoplastic…

Fluid Dynamics · Physics 2026-04-01 Arjun Sharma , Sabarish V. Narayanan , Sarah Hormozi , Donald L. Koch

Bioconvection is the prototypical active matter system for hydrodynamic instabilities and pattern formation in suspensions of biased swimming microorganisms, particularly at the dilute end of the concentration spectrum where cell-cell…

Soft Condensed Matter · Physics 2025-01-22 Martin A. Bees , Prasad Perlekar

Self-propelled phoretic swimmers are generally studied in the laminar flow regime, where their low speed renders inertial effects negligible and trajectories highly predictable. This research tackles the challenge of propulsion in the…

Fluid Dynamics · Physics 2026-01-19 Alessandro Foradori , Paolo Bettotti

The effects of elasticity on filament thinning and breakup are investigated in microchannel cross flow. When a viscous solution is stretched by an external immiscible fluid, a low 100 ppm polymer concentration strongly affects the breakup…

Fluid Dynamics · Physics 2008-04-14 P. E. Arratia , J. P. Gollub , D. J. Durian

We show using theory and experiments that a small particle moving along an elastic membrane through a viscous fluid is repelled from the membrane due to hydro-elastic forces. The viscous stress field produces an elastic disturbance leading…

Soft Condensed Matter · Physics 2018-02-23 Bhargav Rallabandi , Naomi Oppenheimer , Matan Yah Ben Zion , Howard A. Stone

In 1951, G.I. Taylor modeled swimming microorganisms by hypothesizing an infinite sheet in 2D moving in a viscous medium due to a wave passing through it. This simple model not only captured the ability of microorganisms to swim due to the…

Soft Condensed Matter · Physics 2024-10-04 Aditya Jha , Yacine Amarouchene , Thomas Salez

The progressive onset of slip at the wall, which corresponds to a slip length increasing with the solicitation time before reaching a plateau, has been investigated for model viscoelastic polymer solutions, allowing one to vary the longest…

The motility behavior of the nematode Caenorhabditis elegans in polymeric solutions of varying concentrations is systematically investigated in experiments using tracking and velocimetry methods. As the polymer concentration is increased,…

Fluid Dynamics · Physics 2014-05-29 D. A. Gagnon , X. N. Shen , P. E. Arratia

Swimming cells often have to self-propel through fluids displaying non-Newtonian rheology. While past theoretical work seems to indicate that stresses arising from complex fluids should systematically hinder low-Reynolds number locomotion,…

Biological Physics · Physics 2015-06-30 Yi Man , Eric Lauga

The collective motion of dust particles during the mode-coupling induced melting of a two-dimensional plasma crystal is explored in molecular dynamics simulations. The crystal is compressed horizontally by an anisotropic confinement. This…

Plasma Physics · Physics 2015-07-17 I. Laut , C. Räth , S. Zhdanov , V. Nosenko , L. Couëdel , H. M. Thomas

We investigate the dynamics of model microswimmers under confinement, in cylindrical geometries, by means of three dimensional direct numerical calculations with fully resolved hydrodynamics. Such swimmers are known to show collective…

Soft Condensed Matter · Physics 2017-09-26 Norihiro Oyama , John Jairo Molina , Ryoichi Yamamoto

The late-stage phase ordering, in $d=2$ dimensions, of symmetric fluid mixtures violates dynamical scaling. We show however that, even at 50/50 volume fractions, if an asymmetric droplet morphology is initially present then this sustains…

Soft Condensed Matter · Physics 2009-11-07 Alexander J. Wagner , M. E. Cates

The hydrodynamic interactions among bacterial cell bodies, flagella, and surrounding boundaries are essential for understanding bacterial motility in complex environments. In this study, we demonstrate that each slender flagellum can be…

Soft Condensed Matter · Physics 2025-01-07 Baopi Liu , Lu Chen , Ji Zhang

To mimic the motion of biological swimmers in bodily fluids, a novel experimental system of micellar solubilization driven active droplets in a visco-elastic polymeric solution is presented. The visco-elastic nature of the medium,…

Soft Condensed Matter · Physics 2023-05-12 Prateek Dwivedi , Atishay Shrivastava , Naveen Tiwari , Dipin Pillai , Rahul Mangal

The swimming of a two-sphere system and of a three-sphere chain in an incompressible viscous fluid is studied on the basis of simplified equations of motion which take account of both Stokes friction and added mass effects. The analysis is…

Fluid Dynamics · Physics 2017-01-04 B. U. Felderhof

An exact description is provided of an almost spherical fluid vesicle with a fixed area and a fixed enclosed volume locally deformed by external normal forces bringing two nearby points on the surface together symmetrically. The conformal…

Soft Condensed Matter · Physics 2013-04-17 Jemal Guven , Pablo Vázquez-Montejo

Cilia and flagella are hair-like appendages that protrude from the surface of a variety of eukaryotic cells and deform in a wavelike fashion to transport fluids and propel cells. Motivated by the ubiquity of non-Newtonian fluids in biology,…

Fluid Dynamics · Physics 2014-03-19 J. Rodrigo Vélez-Cordero , Eric Lauga

To explore and react to their environment, living micro-swimmers have developed sophisticated strategies for locomotion - in particular, motility with multiple gaits. To understand the physical principles associated with such a behavioural…