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Related papers: Mixing by Swimming Algae

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In the presence of a laminar shear flow, the diffusion of passive colloidal particles is enhanced in the direction parallel to the flow. This classical phenomenon is known as Taylor-Aris dispersion. Besides, microorganisms, such as active…

Many microswimmers are able to swim through viscous fluids by employing periodic non-reciprocal deformations of their appendages. Here we use a simple microswimmer model inspired by swimming biflagellates which consists of a spherical cell…

Soft Condensed Matter · Physics 2025-08-22 Sridhar Bulusu , Andreas Zöttl

It has long been believed that swimming eukaryotes feel solid boundaries through direct ciliary contact. Specifically, based on observations of behavior of green alga Chlamydomonas reinhardtii it has been reported that it is their "flagella…

Fluid Dynamics · Physics 2018-07-18 Mehdi Mirzakhanloo , Mohammad-Reza Alam

Understanding the stochastic dynamics of tracer particles in active fluids is important for identifying the physical properties of flow generating objects such as colloids, bacteria or algae. Here, we study both analytically and numerically…

Soft Condensed Matter · Physics 2010-10-12 Jörn Dunkel , Victor B. Putz , Irwin M. Zaid , Julia M. Yeomans

The measurement of a quantitative and macroscopic parameter to estimate the global motility of a large population of swimming biological cells is a challenge Experiments on the rheology of active suspensions have been performed. Effective…

Soft Condensed Matter · Physics 2014-12-12 Salima Rafai , Levan Jibuti , Philippe Peyla

Swimming cells and microorganisms must often move though complex fluids that contain an immersed microstructure such as polymer molecules, or filaments. In many important biological processes, such as mammalian reproduction and bacterial…

Fluid Dynamics · Physics 2018-08-06 Arshad Kamal , Eric E Keaveny

We present the first time-resolved measurements of the oscillatory velocity field induced by swimming unicellular microorganisms. Confinement of the green alga C. reinhardtii in stabilized thin liquid films allows simultaneous tracking of…

Fluid Dynamics · Physics 2015-05-19 Jeffrey S. Guasto , Karl A. Johnson , J. P. Gollub

Cilia and flagella often exhibit synchronized behavior; this includes phase locking, as seen in {\it Chlamydomonas}, and metachronal wave formation in the respiratory cilia of higher organisms. Since the observations by Gray and Rothschild…

Soft Condensed Matter · Physics 2016-05-19 Kirsty Y. Wan , Raymond E. Goldstein

Cooperation between micro-organisms give rise to novel phenomena like clustering, swarming in suspension. We study the collective behavior of the artificial swimmer called Taylor line at low Reynolds number using multi-particle collision…

Soft Condensed Matter · Physics 2018-02-14 Adyant Agrawal , Sujin B. Babu

Flagella allow eukaryotic cells to move and pump fluid. We present the first three-dimensional, time-resolved imaging of the flagellar waveform of Chlamydomonas reinhardtii, a model alga found in fresh water. During the power stroke, we…

Biological Physics · Physics 2022-10-18 Laurence G. Wilson , Martin A. Bees

We consider two systems of active swimmers moving close to a solid surface, one being a living population of wild-type \textit{E. coli} and the other being an assembly of self-propelled Au-Pt rods. In both situations, we have identified two…

We consider the tracer diffusion $D_{rr}$ that arises from the run-and-tumble motion of low Reynolds number swimmers, such as bacteria. Assuming a dilute suspension, where the bacteria move in uncorrelated runs of length $\lambda$, we…

Soft Condensed Matter · Physics 2015-06-16 Dmitri O. Pushkin , Julia M. Yeomans

Tracer particles immersed in suspensions of biological microswimmers such as E. coli or C. reinhardtii display phenomena unseen in conventional equilibrium systems, including strongly enhanced diffusivity relative to the Brownian value and…

Soft Condensed Matter · Physics 2022-01-27 Henrik Nordanger , Alexander Morozov , Joakim Stenhammar

Enhanced diffusion of passive tracers immersed in active fluids is a universal feature of active fluids and has been extensively studied in recent years. Similar to microrheology for equilibrium complex fluids, the unusual enhanced particle…

Soft Condensed Matter · Physics 2016-11-03 Ou Yang , Yi Peng , Zhengyang Liu , Chao Tang , Xinliang Xu , Xiang Cheng

A dilute non-interacting suspension of micro-swimmers exhibits a finite velocity variance and short-ranged correlations that decay over a swimmer length. For a suspension of interacting straight swimmers, however, pair-interactions leads to…

Soft Condensed Matter · Physics 2020-12-02 Sankalp Nambiar , Piyush Garg , Ganesh Subramanian

Swimming bacteria create long-range velocity fields that stir a large volume of fluid and move around passive particles dispersed in the fluid. Recent experiments and simulations have shown that long-time mean-squared displacement of…

Soft Condensed Matter · Physics 2013-08-20 Alexander Morozov , Davide Marenduzzo

The interaction between swimming microorganisms or artificial self-propelled colloids and passive (tracer) particles in a fluid leads to enhanced diffusion of the tracers. This enhancement has attracted strong interest, as it could lead to…

Soft Condensed Matter · Physics 2017-07-04 Joost de Graaf , Joakim Stenhammar

We analyze the dynamics of a tracer particle embedded in a bath of hard spheres confined in a channel of varying section. By means of Brownian dynamics simulations we apply a constant force on the tracer particle and discuss the dependence…

Soft Condensed Matter · Physics 2018-11-14 Antonio M. Puertas , Paolo Malgaretti , Ignacio Pagonabarraga

Run-and-tumble (RT) motion is commonly observed in flagellated microswimmers, arising from synchronous and asynchronous flagellar beating. One such example is a biflagellated alga, called \textit{Chlamydomonas reinhardtii}. Its flagellar…

Soft Condensed Matter · Physics 2025-10-17 Somnath Paramanick , Umashankar Pardhi , Harsh Soni , Nitin Kumar

Self-propulsion of cellular microswimmers generates flow signatures, commonly classified as pusher- and puller-type, which characterize hydrodynamic interactions with other cells or boundaries. Using experimentally measured beat patterns,…

Cell Behavior · Quantitative Biology 2016-01-20 Gary S. Klindt , Benjamin M. Friedrich