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Regulation of contraction in striated muscle is controlled by a dual mechanism involving both thin filaments containing actin and thick filaments containing myosin. The thin filament is activated by calcium ions binding to troponin, leading…

Mechanically induced folding of passive cross-linkers is a fundamental biological phenomenon. A typical example is a conformational change in myosin II responsible for the power-stroke in skeletal muscles. In this paper we present an…

Biological Physics · Physics 2017-09-14 Matthieu Caruel , Jean-Marc Allain , Lev Truskinovsky

Muscles sense internally generated and externally applied forces, responding to these in a coordinated hierarchical manner at different time scales. The center of the basic unit of the muscle, the sarcomeric M-band, is perfectly placed to…

Tissues and Organs · Quantitative Biology 2021-09-22 Neil Ibata , Eugene M. Terentjev

Myosin II isoforms with varying mechanochemistry and filament size interact with filamentous actin (F-actin) networks to generate contractile forces in cells. How their properties control force production in environments with varying…

Subcellular Processes · Quantitative Biology 2014-07-09 Samantha Stam , Jon Alberts , Margaret L. Gardel , Edwin Munro

We discuss a theoretical model for the cooperative binding dynamics of tropomyosin to actin filaments. Tropomyosin binds to actin by occupying seven consecutive monomers. The model includes a strong attraction between attached tropomyosin…

Soft Condensed Matter · Physics 2009-11-07 Andrej Vilfan

Non-processive molecular motors have to work together in ensembles in order to generate appreciable levels of force or movement. In skeletal muscle, for example, hundreds of myosin II molecules cooperate in thick filaments. In non-muscle…

Biological Physics · Physics 2015-06-16 Thorsten Erdmann , Philipp J. Albert , Ulrich S. Schwarz

In this paper we report, clarify and broaden various recent efforts to complement the chemistry-centered models of force generation in muscles by mechanics-centered models. The physical mechanisms of interest can be grouped into two…

Biological Physics · Physics 2018-06-26 Matthieu Caruel , Lev Truskinovsky

We study the high-frequency micro-mechanical response of suspensions composed by cardiac and skeletal muscle myosin by optical trapping interferometry. We observe that in low ionic strength solutions upon the addition of magnesium adenosine…

Soft Condensed Matter · Physics 2024-10-15 Pablo Domínguez-García , Jose R. Pinto , Ana Akrap , Sylvia Jeney

Actomyosin networks are major structural components of the cell. They provide mechanical integrity and allow dynamic remodeling of eukaryotic cells, self-organizing into the diverse patterns essential for development. We provide a…

Biological Physics · Physics 2012-04-17 Shenshen Wang , Peter G. Wolynes

Cooperative action of molecular motors is essential for many cellular processes. One possible regulator of motor coordination is the elasticity-mediated crosstalk (EMC) coupling between myosin II motors whose origin is the tensile stress…

Biological Physics · Physics 2015-06-15 Nadiv Dharan , Oded Farago

Myosin II plays a pivotal role in muscle contraction by generating force through the cooperative action of multiple motors on actin filaments. In this study, we integrate the nonlinear elasticity of the neck linker in individual myosin II…

Biological Physics · Physics 2023-06-21 Beibei Shen , Yunxin Zhang

Previous models of the cerebrovascular smooth muscle cell have not addressed the interaction between the electrical, chemical and mechanical components of cell function during the development of active tension. These models are primarily…

Cell Behavior · Quantitative Biology 2013-03-27 Jin Yang , John W. Clark , Robert M. Bryan , Claudia S. Robertson

Myosin motor is the machine, which performs mechanical work in the course of adenosine triphosphate molecule hydrolysis and myosin head conformations accompanying this process. For displacement of individual fragments of large molecule…

Biomolecules · Quantitative Biology 2007-05-23 E. V. Rosenfeld

Interactions between a protein and a ligand are often accompanied by a redistribution of the population of thermally accessible conformations. This dynamic response of the protein's functional energy landscape enables a protein to modulate…

Biological Physics · Physics 2016-03-15 Prithviraj Nandigrami , John J. Portman

Contractile forces are essential for many developmental processes involving cell shape change and tissue deformation. Recent experiments on reconstituted actomyosin networks, the major component of the contractile machinery, have shown that…

Biological Physics · Physics 2012-05-31 Shenshen Wang , Peter G. Wolynes

Tissue cells are in a state of permanent mechanical tension that is maintained mainly by myosin II minifilaments, which are bipolar assemblies of tens of myosin II molecular motors contracting actin networks and bundles. Here we introduce a…

Subcellular Processes · Quantitative Biology 2014-09-16 Philipp J. Albert , Thorsten Erdmann , Ulrich S. Schwarz

Cell motility and tissue morphogenesis depend crucially on the dynamic remodelling of actomyosin networks. An actomyosin network consists of an actin polymer network connected by crosslinker proteins and motor protein myosins that generate…

Soft Condensed Matter · Physics 2016-05-25 Chiu Fan Lee , Gunnar Pruessner

Many cell membrane proteins that bind to actin form dynamic clusters driven by contractile flows generated by the actomyosin machinery at the cell cortex. Recent evidence suggests that a necessary condition for the generation of these…

Soft Condensed Matter · Physics 2021-05-25 Sk Raj Hossein , Rituparno Mandal , Madan Rao

Living cells adapt and respond actively to the mechanical properties of their environment. In addition to biochemical mechanotransduction, evidence exists for a myosin-dependent, purely mechanical sensitivity to the stiffness of the…

Myosin II is the muscle molecular motor that works in two bipolar arrays in each thick filament of the striated (skeletal and cardiac) muscle, converting the chemical energy into steady force and shortening by cyclic ATP--driven…

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