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The mechanism of hydrodynamics-induced pairing of soft particles, namely closed bilayer membranes (vesicles, a model system for red blood cells) and drops, is studied numerically with a special attention paid to the role of the confinement…

Gyrotactic algae are bottom heavy, motile cells whose swimming direction is determined by a balance between a buoyancy torque directing them upwards and fluid velocity gradients. Gyrotaxis has, in recent years, become a paradigmatic model…

Chaotic Dynamics · Physics 2019-03-25 Massimo Cencini , Guido Boffetta , Matteo Borgnino , Filippo De Lillo

It has long been believed that swimming eukaryotes feel solid boundaries through direct ciliary contact. Specifically, based on observations of behavior of green alga Chlamydomonas reinhardtii it has been reported that it is their "flagella…

Fluid Dynamics · Physics 2018-07-18 Mehdi Mirzakhanloo , Mohammad-Reza Alam

Self-propelled point-like particles move along circular trajectories when their translocation velocity is constant and the angular velocity related to their orientation vector is also constant. We investigate the collective behavior of…

Biological Physics · Physics 2014-02-04 Yingzi Yang , Feng Qiu , Gerhard Gompper

Motivated by the swimming of sperm in the non-Newtonian fluids of the female mammalian reproductive tract, we examine the swimming of filaments in the nonlinear viscoelastic Upper Convected Maxwell model. We obtain the swimming velocity and…

Soft Condensed Matter · Physics 2009-11-13 Henry C. Fu , Thomas R. Powers , Charles W. Wolgemuth

An interacting pair of chemotactic (anti-chemotactic) active colloids, that can rotate their axes of self-propulsion to align {parallel (anti-parallel)} to a chemical gradient, shows dynamical behaviour that varies from bound states to…

Biological Physics · Physics 2025-10-08 Suropriya Saha , Sriram Ramaswamy , Ramin Golestanian

This study systematically investigates the dynamics of the bacterial transition from free-swimming to surface adhesion, a process characterized by both height $h$ and inclination angle $\Psi$. The surface entrapment process is divided into…

Soft Condensed Matter · Physics 2025-06-25 Baopi Liu , Bowen Jin , Ning An

Most motile bacteria swim in viscous fluids by rotating multiple helical flagellar filaments. These semi-rigid filaments repeatedly join ('bundle') and separate ('unbundle'), resulting in a two-gait random walk-like motion of the cell. In…

Fluid Dynamics · Physics 2020-11-18 Alexander Chamolly , Eric Lauga

Elastic objects across a wide range of scales deform under local changes of their intrinsic properties, yet the shapes are ${\it glocal}$, set by a complicated balance between local properties and global geometric constraints. Here, we…

Soft Condensed Matter · Physics 2025-12-12 Pierre A. Haas , Raymond E. Goldstein

The beating flagella of the green alga Chlamydomonas reinhardtii play a prominent role in cellular mechanics, enabling cells to both displace and sense surrounding fluid. Specifically, flagellum-induced fluid transport enables microalgae to…

Fluid Dynamics · Physics 2026-03-10 Md Iftekhar Yousuf Emon , Gregorius R. Pradipta , Xiang Cheng , Xin Yong

We present laboratory experiments of surface wave turbulence excited by paddles in the deep water regime. The free surface is seeded with buoyant particles that are advected and dispersed by the flow. Positions and velocities of the…

We explore hydrodynamic interactions between microswimmers and corrugated, or rough, surfaces, as found often in biological systems and microfluidic devices. Using the Lorentz reciprocal theorem for viscous flows we derive exact expressions…

Soft Condensed Matter · Physics 2022-08-10 Christina Kurzthaler , Howard A. Stone

We show using theory and experiments that a small particle moving along an elastic membrane through a viscous fluid is repelled from the membrane due to hydro-elastic forces. The viscous stress field produces an elastic disturbance leading…

Soft Condensed Matter · Physics 2018-02-23 Bhargav Rallabandi , Naomi Oppenheimer , Matan Yah Ben Zion , Howard A. Stone

Synchronization of actively oscillating organelles such as cilia and flagella facilitates self-propulsion of cells and pumping fluid in low Reynolds number environments. To understand the key mechanism behind synchronization induced by…

Soft Condensed Matter · Physics 2015-05-20 Nariya Uchida , Ramin Golestanian

We study the stochastic hydrodynamics of colonies of flagellated swimming cells, typified by multicellular choanoflagellates, which can form both rosette and chainlike shapes. The objective is to link cell-scale dynamics to colony-scale…

Quantitative Methods · Quantitative Biology 2023-07-21 Yonatan Ashenafi , Peter R. Kramer

Flagellar-driven locomotion plays a critical role in bacterial attachment and colonization of surfaces, contributing to the risks of contamination and infection. Tremendous attempts to uncover the underlying principles governing bacterial…

Soft Condensed Matter · Physics 2025-02-19 Xin-Xin Xu , Yangguang Tian , Yuhe Pu , Bingchen Che , Hao Luo , Yanan Liu , Yan-Jun Liu , Guangyin Jing

Micro-organisms propel themselves in viscous environments by the periodic, nonreciprocal beating of slender appendages known as flagella. Active materials have been widely exploited to mimic this form of locomotion. However, the realization…

Soft Condensed Matter · Physics 2024-08-06 Ariel Surya Boiardi , Giovanni Noselli

A growing body of work aims at designing and testing micron-scale synthetic swimmers. One method, inspired by the locomotion of flagellated bacteria, consists of applying a rotating magnetic field to a rigid, helically-shaped, propeller…

Fluid Dynamics · Physics 2014-02-17 Yi Man , Eric Lauga

Active particles (living or synthetic) often move through inhomogeneous environments, such as gradients in light, heat or nutrient concentration, that can lead to directed motion (or taxis). Recent research has explored inhomogeneity in the…

Soft Condensed Matter · Physics 2022-08-23 Jiahao Gong , Vaseem A. Shaik , Gwynn J. Elfring

We use a three-bead-spring model to investigate the dynamics of bi-flagellate micro-swimmers near a surface. While the primary dynamics and scattering are governed by geometric-dependent direct contact, the fluid flows generated by the…

Biological Physics · Physics 2017-08-17 Enkeleida Lushi , Vasily Kantsler , Raymond E. Goldstein
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