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Disordered filamentous networks with compliant crosslinks exhibit a low linear elastic shear modulus at small strains, but stiffen dramatically at high strains. Experiments have shown that the elastic modulus can increase by up to three…

Biological Physics · Physics 2015-01-06 Knut M. Heidemann , Abhinav Sharma , Florian Rehfeldt , Christoph F. Schmidt , Max Wardetzky

The semiflexible F-actin network of the cytoskeleton is cross-linked by a variety of proteins including filamin, which contain Ig-domains that unfold under applied tension. We examine a simple semiflexible network model cross-linked by such…

Soft Condensed Matter · Physics 2009-11-11 B. A. DiDonna , Alex J. Levine

Networks of filamentous proteins play a crucial role in cell mechanics. These cytoskeletal networks, together with various crosslinking and other associated proteins largely determine the (visco)elastic response of cells. In this letter we…

Disordered Systems and Neural Networks · Physics 2009-11-10 D. A. Head , A. J. Levine , F. C. MacKintosh

Motivated by recent experiments showing that stiff biopolymer gels exhibit highly unusual negative normal elastic stresses, we develop a computational model for stiff polymer networks subject to large strains. In all cases, we find that…

Soft Condensed Matter · Physics 2009-11-13 Enrico Conti , F. C. MacKintosh

We investigate theoretically the effect of polymer tension on the collective behavior of reversibly binding cross-links. For this purpose, we employ a model of two weakly bending wormlike chains aligned in parallel by a tensile force, with…

Biological Physics · Physics 2015-02-10 Panayotis Benetatos , Alice von der Heydt , Annette Zippelius

The collective action of actively contractile units embedded in elastic biopolymer networks plays a crucial role in regulating the network's macroscopic mechanical response. Here, we investigate how the macroscopic boundary stress in model…

Soft Condensed Matter · Physics 2026-03-25 Abhinav Kumar , David A. Quint , Kinjal Dasbiswas

Networks of semiflexible or stiff polymers such as most biopolymers are known to deform inhomogeneously when sheared. The effects of such non-affine deformation have been shown to be much stronger than for flexible polymers. To date, our…

Soft Condensed Matter · Physics 2023-03-08 Sihan Chen , Tomer Markovich , Fred C. MacKintosh

We review the physical properties of macromolecular networks, consisting of semiflexible polymers such as actin. We start by giving a theoretical analysis of the conformational statistics and mechanical response of single filaments.…

Soft Condensed Matter · Physics 2008-02-03 Erwin Frey , Klaus Kroy , Jan Wilhelm , Erich Sackmann

Stretching an elastic material along one axis typically induces contraction along the transverse axes, a phenomenon known as the Poisson effect. From these strains, one can compute the specific volume, which generally either increases or,…

Soft Condensed Matter · Physics 2024-07-31 Jordan L. Shivers , Fred C. MacKintosh

We use optical tweezers microrheology and fluorescence microscopy to apply nonlinear microscale strains to entangled and crosslinked actin networks, and measure the resulting stress and actin filament deformations. We couple nonlinear…

Soft Condensed Matter · Physics 2016-10-24 Bekele Gurmessa , Shea Ricketts , Rae M. Robertson-Anderson

We study the mechanical stiffening behavior in two-dimensional (2D) cross-linked networks of semiflexible biopolymer filaments under simple shear. Filamental constituents immersed in a fluid undergo thermally excited bending motions.…

Biological Physics · Physics 2007-05-23 T. Van Dillen , P. R. Onck , E. Van der Giessen

We propose a class of microstructurally informed models for the linear elastic mechanical behavior of cross-linked polymer networks such as the actin cytoskeleton. Salient features of the models include the possibility to represent…

Other Quantitative Biology · Quantitative Biology 2008-02-27 Ronald Y. Kwon , Adrian J. Lew , Christopher R. Jacobs

Cells make use of semi-flexible biopolymers such as actin or intermediate filaments to control their local viscoelastic response by dynamically adjusting the concentration and type of cross-linker molecules. The microstructure of the…

Soft Condensed Matter · Physics 2009-11-13 Yuxia Luan , Oliver Lieleg , Bernd Wagner , Andreas R. Bausch

Transient networks comprised of polymers connected by short-lived bonds are a common design theme for both biological and synthetic materials. Transient bonds can provide mechanical rigidity, while still allowing for visco-elastic flows on…

Soft Condensed Matter · Physics 2019-01-02 Yuval Mulla , Gijsje H. Koenderink

We develop a continuum theory for equilibrium elasticity of a network of crosslinked semiflexible filaments, spanning the full range between flexible entropy-driven chains to stiff athermal rods. We choose the 3-chain constitutive model…

Soft Condensed Matter · Physics 2016-07-12 Fanlong Meng , Eugene M. Terentjev

We formulate an effective medium (mean field) theory of a material consisting of randomly distributed nodes connected by straight slender rods, hinged at the nodes. Defining novel wavelength-dependent effective elastic moduli, we calculate…

Soft Condensed Matter · Physics 2015-06-05 J. I. Katz , J. J. Hoffman , M. S. Conradi , J. G. Miller

While polymer solutions lack the mechanical stability only transiently cross-linked networks can fulfill the competing requirements of structural stability and maximal energy dissipation. Here, we show that transient cross-links entail…

Soft Condensed Matter · Physics 2009-11-13 O. Lieleg , M. M. A. E. Claessens , Y. Luan , A. R. Bausch

Disordered spring networks can exhibit rigidity transitions, due to either the removal of materials in over-constrained networks or the application of strain in under-constrained ones. While an effective medium theory (EMT) exists for the…

Soft Condensed Matter · Physics 2022-04-04 Ojan Khatib Damavandi , M. Lisa Manning , J. M. Schwarz

We report on the nonlinear mechanical properties of a statistically homogeneous, isotropic semiflexible network cross-linked by polymers containing numerous small unfolding domains, such as the ubiquitous F-actin cross-linker Filamin. We…

Soft Condensed Matter · Physics 2013-05-29 B. A. DiDonna , Alex J. Levine

Soft living tissues like cartilage can be considered as biphasic materials comprised of a fibrous complex biopolymer network and a viscous background liquid. Here, we show by a combination of experiment and theoretical analysis that both…

Soft Condensed Matter · Physics 2019-10-02 Melle T. J. J. M. Punter , Bart E. Vos , Bela M. Mulder , Gijsje H. Koenderink