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In a world without inertia, Purcell's scallop theorem states that in a Newtonian fluid a time-reversible motion cannot produce any net force or net flow. Here we consider the extent to which the nonlinear rheological behavior of…

Fluid Dynamics · Physics 2010-04-09 On Shun Pak , Eric Lauga

Fluid-suspended microorganisms have evolved different swimming and feeding strategies in order to cope with an environment dominated by viscous effects. For instance ciliated organisms rely on the collective motion of flexible appendices to…

Fluid Dynamics · Physics 2015-06-19 Roger M. Arco , J. Rodrigo Velez-Cordero , Eric Lauga , Roberto Zenit

In isotropic fluids like water, micrometer-scale swimmers have evolved swim strokes to translate despite their tiny size. As described by Purcell in his Scallop Theorem, reciprocal motions, like those performed by a scallop, cannot drive…

Many microorganisms swim through gels and non-Newtonian fluids in their natural environments. In this paper, we focus on microorganisms which use flagella for propulsion. We address how swimming velocities are affected in nonlinearly…

Biological Physics · Physics 2010-04-07 Henry C. Fu , Charles W. Wolgemuth , Thomas R. Powers

Understanding swimming in soft yielding media is challenging due to their complex deformation response to the swimmer's motion. We experimentally show that a scallop-inspired swimmer with reciprocally flapping wings generates locomotion in…

Soft Condensed Matter · Physics 2024-12-09 Hongyi Xiao , Harol Torres , Achim Sack , Thorsten Pöschel

Actuating periodically an elastic filament in a viscous liquid generally breaks the constraints of Purcell's scallop theorem, resulting in the generation of a net propulsive force. This observation suggests a method to design simple…

Soft Condensed Matter · Physics 2009-09-29 Eric Lauga

When an elastic object is dragged through a viscous fluid tangent to a rigid boundary, it experiences a lift force perpendicular to its direction of motion. An analogous lift mechanism occurs when a rigid symmetric object translates…

Fluid Dynamics · Physics 2018-08-22 Abdallah Daddi-Moussa-Ider , Bhargav Rallabandi , Stephan Gekle , Howard A. Stone

Many cells exploit the bending or rotation of flagellar filaments in order to self-propel in viscous fluids. While appropriate theoretical modelling is available to capture flagella locomotion in simple, Newtonian fluids, formidable…

Biological Physics · Physics 2017-08-02 Emily E. Riley , Eric Lauga

Purcell's scallop theorem defines the type of motions of a solid body - reciprocal motions - which cannot propel the body in a viscous fluid with zero Reynolds number. For example, the flapping of a wing is reciprocal and, as was recently…

Soft Condensed Matter · Physics 2008-10-02 Eric Lauga

Biological locomotion in nature is often achieved by the interaction between a flexible body and its surrounding medium. The interaction of a flexible body with granular media is less understood compared with viscous fluids partially due to…

Fluid Dynamics · Physics 2017-08-03 Zhiwei Peng , Yang Ding , Kyle Pietrzyk , Gwynn J. Elfring , On Shun Pak

We experimentally study a scallop-like swimmer with reciprocally flapping wings in a nearly frictionless, cohesive granular medium consisting of hydrogel spheres. Significant locomotion is found when the swimmer's flapping frequency matches…

Soft Condensed Matter · Physics 2026-04-29 Hongyi Xiao , Jing Wang , Achim Sack , Ralf Stannarius , Thorsten Pöschel

In this work we mimic the efficient propulsion mechanism of natural cilia by magnetically actuating thin films in a cyclic but non-reciprocating manner. By simultaneously solving the elasto-dynamic, magnetostatic and fluid mechanics…

Conventionally, a microscopic particle that performs a reciprocal stroke cannot move through its environment. This is because at small scales, the response of simple Newtonian fluids is purely viscous and flows are time-reversible. We show…

Fluid Dynamics · Physics 2012-08-21 Nathan C. Keim , Mike Garcia , Paulo E. Arratia

Thin cylindrical membranes arise in a wide variety of biological systems ranging from tubular structures on and within cell membranes to in-vitro experiments on artificial vesicles. Motor proteins embedded in such fluidic membranes often…

Fluid Dynamics · Physics 2025-07-09 Udaya Maurya , Surya Teja Gavva , Arpan Saha , Rickmoy Samanta

Locomotion on small scales is dominated by the effects of viscous forces and, as a result, is subject to strong physical and mathematical constraints. Following Purcell's statement of the scallop theorem which delimitates the types of…

Biological Physics · Physics 2011-08-30 Eric Lauga

Micropolar fluid theory, an extension of classical Newtonian fluid dynamics, incorporates angular velocities and rotational inertias and has long been a foundational framework for describing granular flows. We propose a macroscopic model of…

Soft Condensed Matter · Physics 2025-12-01 Jarosław Pawłowski , Marcin Dudziak , Matteo Baggioli , Jie Zhang , Piotr Surówka

We construct viscous fluid flow sourced by a force dipole embedded in a cylindrical fluid membrane, coupled to external embedding fluids. We find analytic expressions for the flow, in the limit of infinitely long and thin tubular membranes.…

Soft Condensed Matter · Physics 2025-06-23 Samyak Jain , Rickmoy Samanta

We use particle simulations to reveal two distinct propulsion mechanisms for a scallop-like swimmer to locomote itself in granular media by reciprocally flapping its wings. Based on the discrete element method, we examine the kinematics and…

Soft Condensed Matter · Physics 2025-10-28 Amir Nazemi , Hongyi Xiao

Natural cilia are hair-like microtubule-based structures that are able to move fluid at low Reynolds number through asymmetric motion. In this paper we follow a biomimetic approach to design artificial cilia lining the inner surface of…

Swimming fish and flying insects use the flapping of fins and wings to generate thrust. In contrast, microscopic organisms typically deform their appendages in a wavelike fashion. Since a flapping motion with two degrees of freedom is able,…

Fluid Dynamics · Physics 2014-06-18 Loic Was , Eric Lauga
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