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The interaction of surfaces in relative motion in wet environments is dominated by lubrication forces, which play a pivotal role in the dynamics of microscopic systems. Here, we develop motile vesicles that exploit lubrication forces to…

We investigate the sliding of objects on an inclined granular surface close to the avalanche threshold. Our experiments show that the stability is driven by the surface deformations. Heavy objects generate footprint-like deformations which…

Soft Condensed Matter · Physics 2017-08-09 Jérôme Crassous , Antoine Humeau , Samuel Boury , Jérôme Casas

We optimize three-dimensional snake kinematics for locomotor efficiency. We assume a general space-curve representation of the snake backbone with small-to-moderate lifting off the ground and negligible body inertia. The cost of locomotion…

Biological Physics · Physics 2022-05-11 Silas Alben

Limbless terrestrial animals exhibit exceptional locomotor versatility and control, currently unmatched by engineered counterparts. Here, we introduce a computational framework that enables soft synthetic snakes to navigate unstructured,…

Robotics · Computer Science 2026-05-26 Xiaotian Zhang , Ali Albazroun , Tixian Wang , Songyuan Cui , Prashant G. Mehta , Mattia Gazzola

We present in this fluid dynamics video a novel experimental setup with self-propelled swimmers on a free surface. The swimmers, modeled as flexible thin filaments, are subjected to external electromagnetic forcing driving a propagating…

Fluid Dynamics · Physics 2014-11-04 Sophie Ramananarivo , Benjamin Thiria , Ramiro Godoy-Diana

A wide range of microorganisms, e.g. bacteria, propel themselves by rotation of soft helical tails, also known as flagella. Due to the small size of these organisms, viscous forces overwhelm inertial effects and the flow is at low Reynolds…

Robotics · Computer Science 2021-03-11 Yayun Du , Andrew Miller , Mohammad Khalid Jawed

The transport of small quantities of liquid on a solid surface is inhibited by the resistance to motion caused by the contact between the liquid and the solid. To overcome such resistance, motion can be externally driven through gradients…

We investigate the self-locomotion of an elongated microswimmer by virtue of the unidirectional tangential surface treadmilling. We show that the propulsion could be almost frictionless, as the microswimmer is propelled forward with the…

Fluid Dynamics · Physics 2009-11-13 Alexander M. Leshansky , Oded Kenneth , Omri Gat , Joseph E. Avron

Undulatory locomotion, a gait in which thrust is produced in the opposite direction of a traveling wave of body bending, is a common mode of propulsion used by animals in fluids, on land, and even within sand. As such it has been an…

Biological Physics · Physics 2013-06-11 Yang Ding , Sarah S. Sharpe , Kurt Wiesenfeld , Daniel I. Goldman

Snake moves across various terrains by bending its elongated body. Recent studies discovered that snakes can use vertical bending to traverse terrain of large height variation, such as horizontally oriented cylinders, a wedge (Jurestovsky,…

Quantitative Methods · Quantitative Biology 2025-09-29 Yifeng Zhang , Qihan Xuan , Qiyuan Fu , Chen Li

Fish swim with flexible fins that stand in stark contrast to the rigid propulsors of engineered vehicles. Using numerical simulations of the dynamics of flow-structure interaction, we have found that dorso-ventral deformation in flexible…

Fluid Dynamics · Physics 2026-04-16 Sushrut Kumar , Matthew J. McHenry , Jung-Hee Seo , Rajat Mittal

Self-propelling organisms locomote via generation of patterns of self-deformation. Despite the diversity of body plans, internal actuation schemes and environments in limbless vertebrates and invertebrates, such organisms often use similar…

A flexible membrane deforming its shape in time can self-propel in a viscous fluid. Alternatively, if the membrane is anchored, its deformation will lead to fluid transport. Past work in this area focused on situations where the deformation…

Soft Condensed Matter · Physics 2013-02-12 Arthur A. Evans , Eric Lauga

Many parts of biological organisms are comprised of deformable porous media. The biological media is both pliable enough to deform in response to an outside force and can deform by itself using the work of an embedded muscle. For example,…

Fluid Dynamics · Physics 2021-08-18 Tagir Farkhutdinov , François Gay-Balmaz , Vakhtang Putkaradze

The intricate wobbling motion of flagellated bacteria, characterized by the periodic precession of the cell body, is a determinant factor in their motility and navigation within complex fluid environments. While well-studied in quiescent…

Soft Condensed Matter · Physics 2026-01-26 Wei Feng , Fanglong Dang , Hao Luo , Alan C. H. Tsang , Yanan Liu , Guangyin Jing

Vesicles are soft elastic bodies with distinctive mechanical properties such as bending resistance, membrane fluidity, and their strong ability to deform, mimicking some properties of biological cells. While previous three-dimensional (3D)…

Soft Condensed Matter · Physics 2021-06-18 Jinming Lyu , Paul G. Chen , Alexander Farutin , Marc Jaeger , Chaouqi Misbah , and Marc Leonetti

In natural environments, solid surfaces present both opportunities and challenges for bacteria. On one hand, they serve as platforms for biofilm formation, crucial for bacterial colonization and resilience in harsh conditions. On the other…

Biological Physics · Physics 2025-07-01 Antai Tao , Guangzhe Liu , Rongjing Zhang , Junhua Yuan

Snakes' bodies are covered in scales that make it easier to slide in some directions than in others. This frictional anisotropy allows for sliding locomotion with an undulatory gait, one of the most common for snakes. Isotropic friction is…

Biological Physics · Physics 2019-06-12 Silas Alben

The ciliary locomotion and feeding of an axisymmetric micro-swimmer in a complex fluid whose viscosity depends on nutrient concentration are investigated numerically. The micro-swimmer is modeled as having spheroidal geometry, and ciliary…

Fluid Dynamics · Physics 2020-07-01 Patrick S. Eastham , Kourosh Shoele

Biological and artificial microswimmers often have to propel through a variety of environments, ranging from heterogeneous suspending media to strong geometrical confinement. Under confinement, local flow fields generated by microswimmers,…

Soft Condensed Matter · Physics 2024-06-04 Florian A. Overberg , Gerhard Gompper , Dmitry A. Fedosov