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Random packings of granular chains are presented as a model polymer system to investigate the contribution of entanglements to strain-stiffening in the absence of Brownian motion. The chain packings are sheared in triaxial compression…

Soft Condensed Matter · Physics 2015-05-30 Eric Brown , Alice Nasto , Athanasios G. Athanassiadis , Heinrich M. Jaeger

We introduce a phenomenological model to study the properties of bundles of polar filaments which interact via active elements. The stability of the homogeneous state, the attractors of the dynamics in the unstable regime and the tensile…

Statistical Mechanics · Physics 2009-10-31 Karsten Kruse , Frank Julicher

Motivated by recent experiments showing that stiff biopolymer gels exhibit highly unusual negative normal elastic stresses, we develop a computational model for stiff polymer networks subject to large strains. In all cases, we find that…

Soft Condensed Matter · Physics 2009-11-13 Enrico Conti , F. C. MacKintosh

Collective dynamics and force generation by cytoskeletal filaments are crucial in many cellular processes. Investigating growth dynamics of a bundle of N independent cytoskeletal filaments pushing against a wall, we show that chemical…

Biological Physics · Physics 2015-06-17 Dipjyoti Das , Dibyendu Das , Ranjith Padinhateeri

We explore the effect of random permanent cross-links on a system of directed polymers confined between two planes with their end-points free to slide on them. We treat the cross-links as quenched disorder and we use a semimicroscopic…

Soft Condensed Matter · Physics 2010-02-22 Stephan Ulrich , Annette Zippelius , Panayotis Benetatos

The growth of actin filament networks is a fundamental biological process that drives a variety of cellular and intracellular motions. During motility, eukaryotic cells and intracellular pathogens are propelled by actin networks organized…

Biological Physics · Physics 2009-11-13 Joshua W. Shaevitz , Daniel A. Fletcher

Self organization mechanisms are essential for the cytoskeleton to adapt to the requirements of living cells. They rely on the intricate interplay of cytoskeletal filaments, crosslinking proteins and molecular motors. Here we present an in…

Biological Physics · Physics 2015-05-28 Simone Köhler , Volker Schaller , Andreas R. Bausch

In cells, cytoskeletal filament networks are responsible for cell movement, growth, and division. Filaments in the cytoskeleton are driven and organized by crosslinking molecular motors. In reconstituted cytoskeletal systems, motor activity…

We optically drive a microsphere at constant speed through entangled actin networks of 0.2 - 1.4 mg/ml at rates faster than the critical rate controlling the onset of a nonlinear response. By measuring the resistive force exerted on the…

Soft Condensed Matter · Physics 2016-09-28 Bekele Gurmessa , Robert Fitzpatrick , Tobias T. Falzone , Rae M. Robertson-Anderson

Filopodia are long, finger-like membrane tubes supported by cytoskeletal filaments. Their shape is determined by the stiffness of the actin filament bundles found inside them and by the interplay between the surface tension and bending…

Subcellular Processes · Quantitative Biology 2008-06-24 Sander Pronk , Phillip L. Geissler , Daniel A. Fletcher

Mutual uncrossability of polymers generates topological constraints on their conformations and dynamics, which are generally described using the tube model. We imaged confinement tubes for individual polymers within a F-actin solution by…

Soft Condensed Matter · Physics 2009-08-20 M. Romanowska , H. Hinsch , N. Kirchgessner , M. Giesen , M. Degawa , B. Hoffmann , E. Frey , R. Merkel

Depletion interactions are thought to significantly contribute to the organization of intracellular structures in the crowded cytosol. The strength of depletion interactions depends on physical parameters like the depletant number density…

In nature interactions between biopolymers and motor proteins give rise to biologically essential emergent behaviours. Besides cytoskeleton mechanics, active nematics arise from such interactions. Here we present a study on 3D active…

The mechanical properties of the cell depend crucially on the tension of its cytoskeleton, a biopolymer network that is put under stress by active motor proteins. While the fibrous nature of the network is known to strongly affect the…

Soft Condensed Matter · Physics 2018-05-15 Pierre Ronceray , Chase P. Broedersz , Martin Lenz

The cytoskeleton is a model active matter system that controls diverse cellular processes from division to motility. While both active actomyosin dynamics and actin-microtubule interactions are key to the cytoskeleton's versatility and…

Pattern formation and the mechanics of a mixture of actin filaments and myosin motors that is confined by a rigid membrane is investigated. By using a coarse-grained molecular dynamics model, we demonstrate that the competition between the…

Soft Condensed Matter · Physics 2023-04-12 Mitsusuke Tarama , Tatsuo Shibata

Cells moving on a two dimensional substrate generate motion by polymerizing actin filament networks inside a flat membrane protrusion. New filaments are generated by branching off existing ones, giving rise to branched network structures.…

Biological Physics · Physics 2015-06-29 Mohammadhosein Razbin , Martin Falcke , Panayotis Benetatos , Annette Zippelius

Actin is one of the most studied cytoskeleton proteins showing a very rich span of structures. It can self-assemble actively into dynamical structures that govern the mechanical properties of the cell, its motility and its division.…

Soft Condensed Matter · Physics 2020-03-17 Maayan Levin , Raya Sorkin , David Pine , Rony Granek , Anne Bernheim-Groswasser , Yael Roichman

Here, the general design and properties of new multiple network elastomers with an exceptional combination of stiffness, toughness, and elasticity are reported. In this paper, it is reported in more detail how the increase in strain at…

Soft Condensed Matter · Physics 2017-02-14 Etienne Ducrot , Costantino Creton

Any macroscopic deformation of a filamentous bundle is necessarily accompanied by local sliding and/or stretching of the constituent filaments. Yet the nature of the sliding friction between two aligned filaments interacting through…