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The macromechanical properties of purely bundled in vitro actin networks are not only determined by the micromechanical properties of individual bundles but also by molecular unbinding events of the actin binding protein (ABP) fascin. Under…

Soft Condensed Matter · Physics 2009-11-13 Oliver Lieleg , Andreas R. Bausch

Recent experiments have demonstrated that the nonlinear elasticity of in vitro networks of the biopolymer actin is dramatically altered in the presence of a flexible cross-linker such as the abundant cytoskeletal protein filamin. The basic…

Soft Condensed Matter · Physics 2015-05-13 C. P. Broedersz , C. Storm , F. C. MacKintosh

Filopodia are bundles of actin filaments that extend out ahead of the leading edge of a crawling cell to probe its upcoming environment. {\it In vitro} experiments [D. Vignjevic {\it et al.}, J. Cell Biol. {\bf 160}, 951 (2003)] have…

Subcellular Processes · Quantitative Biology 2010-05-14 K. -C. Lee , A. Gopinathan , J. M. Schwarz

Actomyosin contractility is essential for biological force generation, and is well understood in highly organized structures such as striated muscle. Additionally, actomyosin bundles devoid of this organization are known to contract both in…

Biological Physics · Physics 2012-04-03 Martin Lenz , Margaret L. Gardel , Aaron R. Dinner

We discuss the response of biopolymer filament bundles bound by transient cross linkers to compressive loading. These systems admit a mechanical instability at stresses typically below that of traditional Euler buckling. In this…

Soft Condensed Matter · Physics 2021-10-13 Valentin M Slepukhin , Alex J Levine

Motor proteins drive persistent motion and self-organisation of cytoskeletal filaments. However, state-of-the-art microscopy techniques and continuum modelling approaches focus on large length and time scales. Here, we perform…

Soft Condensed Matter · Physics 2024-06-03 Gerrit Vliegenthart , Arvind Ravichandran , Marisol Ripoll , Thorsten Auth , Gerhard Gompper

Essentially all biology is active and dynamic. Biological entities autonomously sense, com- pute, and respond using energy-coupled ratchets that can produce force and do work. The cytoskeleton, along with its associated proteins and motors,…

Soft Condensed Matter · Physics 2017-03-28 Kasimira T. Stanhope , Vikrant Yadav , Christian D. Santangelo , Jennifer L. Ross

Living cells move thanks to assemblies of actin filaments and myosin motors that range from very organized striated muscle tissue to disordered intracellular bundles. The mechanisms powering these disordered structures are debated, and all…

Biological Physics · Physics 2017-09-26 Martin Lenz

Motivated by recent experiments showing nonlinear elasticity of in vitro networks of the biopolymer actin cross-linked with filamin, we present an effective medium theory of flexibly cross-linked stiff polymer networks. We model such…

Soft Condensed Matter · Physics 2008-10-19 C. P. Broedersz , C. Storm , F. C. MacKintosh

Topological defects are crucial to the thermodynamics and structure of condensed matter systems. For instance, when incorporated into crystalline membranes like graphene, disclinations with positive and negative topological charge…

Soft Condensed Matter · Physics 2018-08-29 Isaac R. Bruss , Gregory M. Grason

To study the compressional forces exerted by a bundle of living stiff filaments pressing on a surface, akin to the case of an actin bundle in filopodia structures, we have performed particulate Molecular Dynamics simulations of a grafted…

Soft Condensed Matter · Physics 2015-06-17 Sanoop Ramachandran , Jean-Paul Ryckaert

Aggregates of twisted protein fibers, such as sickle hemoglobin and actin, are important examples of biopolymers in which elastic interactions play a crucial role in determining the (metastable) bundle radii. Here, we present a corrected…

Soft Condensed Matter · Physics 2010-02-15 Y. Liu , J. D. Gunton

Understanding the role of non-equilibrium driving in self-organization is crucial for developing a predictive description of biological systems, yet it is impeded by their complexity. The actin cytoskeleton serves as a paradigm for how…

Soft Condensed Matter · Physics 2020-12-22 Yuqing Qiu , Michael Nguyen , Glen M. Hocky , Aaron R. Dinner , Suriyanarayanan Vaikuntanathan

Cross-linked polymer networks with orientational order constitute a wide class of soft materials and are relevant to biological systems (e.g., F-actin bundles). We analytically study the nonlinear force-extension relation of an array of…

Disordered Systems and Neural Networks · Physics 2015-06-05 Panayotis Benetatos , Stephan Ulrich , Annette Zippelius

Densely-packed bundles of biological filaments (filamentous proteins) are common and critical structural elements in range of biological materials. While most bundles form from intrinsically straight filaments, there are notable examples of…

Soft Condensed Matter · Physics 2013-01-23 Gregory M. Grason

Structure formation and constant reorganization of the actin cytoskeleton are key requirements for the function of living cells. Here we show that a minimal reconstituted system consisting of actin filaments, crosslinking molecules and…

Biological Physics · Physics 2015-05-27 Simone Köhler , Volker Schaller , Andreas R. Bausch

Disordered networks of semiflexible filaments are common support structures in biology. Familiar examples include fibrous matrices in blood clots, bacterial biofilms, and essential components of cells and tissues of plants, animals, and…

Soft Condensed Matter · Physics 2023-09-28 Saamiya Syed , Fred C. MacKintosh , Jordan L. Shivers

We present a simple model for the bidirectional dynamics of actin bundles with alternating polarities in gliding assays with non-processive myosin motors. In the model, the bundle is represented as an elastic chain consisting of monomers…

Soft Condensed Matter · Physics 2015-05-19 Barak Gur , Oded Farago

We examine the nonequilibrium production of topological defects -- braids -- in semiflexible filament bundles under cycles of compression and tension. During these cycles, the period of compression facilitates the thermally activated pair…

Soft Condensed Matter · Physics 2021-05-28 Valentin M. Slepukhin , Alex J. Levine

The crystallization of bundles in filament networks interacting via long-range repulsions in confinement is described by a phenomenological model. The model demonstrates the formation of the hexagonal crystalline order via the interplay of…

Soft Condensed Matter · Physics 2013-05-10 Zhenwei Yao , Monica Olvera de la Cruz