English

Travelling waves are hydrodynamically optimal for long-wavelength flagella

Fluid Dynamics 2020-11-18 v1 Soft Condensed Matter Biological Physics

Abstract

Swimming eukaryotic microorganisms such as spermatozoa, algae and ciliates self-propel in viscous fluids using travelling wave-like deformations of slender appendages called flagella. Waves are predominant because Purcell's scallop theorem precludes time-reversible kinematics for locomotion. Using the calculus of variations on a periodic long-wavelength model of flagellar swimming, we show that the planar flagellar kinematics maximising the time-averaged propulsive force for a fixed amount of energy dissipated in the surrounding fluid correspond for all times to waves travelling with constant speed, potentially on a curved centreline, with propulsion always in the direction opposite to the wave.

Keywords

Cite

@article{arxiv.2011.08778,
  title  = {Travelling waves are hydrodynamically optimal for long-wavelength flagella},
  author = {Eric Lauga},
  journal= {arXiv preprint arXiv:2011.08778},
  year   = {2020}
}
R2 v1 2026-06-23T20:19:19.946Z