Lagrangian 4-planes in holomorphic symplectic varieties of K3^[4] type

We classify the cohomology classes of Lagrangian 4-planes $\P^4$ in a smooth manifold $X$ deformation equivalent to a Hilbert scheme of 4 points on a $K3$ surface, up to the monodromy action. Classically, the cone of effective curves on a $K3$ surface $S$ is generated by nonegative classes $C$, for which $(C,C)\geq0$, and nodal classes $C$, for which $(C,C)=-2$; Hassett and Tschinkel conjecture that the cone of effective curves on a holomorphic symplectic variety $X$ is similarly controlled by "nodal" classes $C$ such that $(C,C)=-\gamma$, for $(\cdot,\cdot)$ now the Beauville-Bogomolov form, where $\gamma$ classifies the geometry of the extremal contraction associated to $C$. In particular, they conjecture that for $X$ deformation equivalent to a Hilbert scheme of $n$ points on a $K3$ surface, the class $C=\ell$ of a line in a smooth Lagrangian $n$-plane $\P^n$ must satisfy $(\ell,\ell)=-\frac{n+3}{2}$. We prove the conjecture for $n=4$ by computing the ring of monodromy invariants on $X$, and showing there is a unique monodromy orbit of Lagrangian 4-planes.